Wallowa beardtongue, Wallowa penstemon
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Blue Mountain penstemon, elegant beardtongue, elegant penstemon, lovely beardtongue, Venus penstemon
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Herbs. |
Subshrubs. |
ascending to erect, 6–26 cm, retrorsely hairy or glabrate, not glaucous. |
ascending to erect, 18–80 cm, glabrous or retrorsely hairy, hairs pointed, not glaucous. |
basal and cauline, ± leathery or not, glabrous or puberulent proximally on petiole and along midvein abaxially; basal and proximal cauline 10–60 × 3–18 mm, blade spatulate to elliptic, base tapered, margins entire, sometimes ± serrulate distally, apex rounded to obtuse or acute; cauline 1–3 pairs, sessile, 14–35 × 2–8 mm, blade lanceolate, sometimes oblanceolate, base truncate, margins entire, rarely ± serrulate distally, apex obtuse to acute. |
cauline, opposite, glabrous, sometimes glaucous; cauline 7–15 pairs, short-petiolate or sessile, (20–)40–100(–120) × (7–)10–30(–38) mm, blade ovate to lanceolate, base tapered to truncate, margins subentire or serrate, apex acute. |
interrupted, cylindric, 1–13 cm, axis moderately glandular-pubescent, verticillasters 1–5, cymes 2–5-flowered, 2 per node; proximal bracts lanceolate, 8–30 × 2–7 mm, margins entire; peduncles and pedicels ascending to erect, glandular-pubescent. |
interrupted, ± secund, (2–)7–40 cm, axis glabrous or retrorsely hairy, verticillasters (3–)5–14, cymes 2–7-flowered, 2 per node; proximal bracts lanceolate, (6–)23–70(–115) × (1–)4–28(–35) mm; peduncles and pedicels ascending to erect, glabrous or retrorsely hairy. |
calyx lobes ovate to lanceolate, 2.6–4.8 × 1–1.8 mm, glandular-pubescent; corolla light blue to light purple, with violet nectar guides, funnelform, 10–14 mm, glandular-pubescent externally, moderately white- or yellowish lanate internally abaxially, tube 4–5 mm, throat slightly inflated, 3.5–4.5 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular to subexplanate, 0.6–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 6–7 mm, included, 0.3–0.4 mm diam., tip recurved, distal 0.5–1.5 mm densely pilose, hairs yellow, to 0.7 mm; style 8–10 mm. |
calyx lobes ovate to obovate or lanceolate, 2.5–6 × 1.3–1.8 mm, glabrous except for margins sometimes ciliate; corolla lavender to purple or violet, with reddish purple nectar guides, weakly ventricose, 24–38 mm, glabrous externally except lobe margins densely white-ciliate, glabrous internally, tube 7–9 mm, throat gradually inflated, 9–10 mm diam., rounded abaxially; stamens: longer pair reaching orifice or exserted, filaments white-lanate distally, rarely glabrous, pollen sacs parallel, 1.6–2.1 mm, distal 1/2–2/3 indehiscent, sides hispidulous, hairs white, to 0.2 mm near point of attachment of connective, sutures denticulate, teeth to 0.2 mm; staminode 16–19 mm, reaching orifice or barely exserted, 0.4–0.5 mm diam., distal 2–5 mm sparsely to moderately lanate, hairs white, to 2 mm; style 18–25 mm. |
4–6 × 2.5–4 mm, glabrous. |
6–9 × 4–5 mm. |
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= 64. |
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Flowering Jul–Aug. |
Flowering (May–)Jun–Sep. |
Open rocky slopes, meadows. |
Rock outcrops, rocky and talus slopes, especially on basalt, shrublands, forest clearings. |
2300–2500 m. (7500–8200 ft.) |
300–2700 m. (1000–8900 ft.) |
OR
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CA; ID; OR; UT; WA
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Penstemon spatulatus is known only from the Strawberry and Wallowa mountains of northeastern Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon venustus is known from the Snake River Plateau in west-central Idaho, northeastern Oregon, and southeastern Washington and is naturalized in Lassen County, California. E. C. Neese and N. D. Atwood (2003) reported P. venustus, apparently introduced, from Duchesne County, Utah. E. Heitz (1927) reported a chromosome number of 2n = 14–16 for P. venustus; that count conflicts with all other reports for P. venustus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 222. |
FNA vol. 17, p. 245. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
Pennell: Notul. Nat. Acad. Nat. Sci. Philadelphia 71: 10. (1941) |
Douglas ex Lindley: Edwards’s Bot. Reg. 16: plate 1309. (1830) — (as Pentstemon venustum) |
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