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side-bells beardtongue, sidebells penstemon

Garrett's beardtongue, White River beardtongue

Habit Plants: caudex sometimes herbaceous.
Stems

ascending to erect, (15–)20–45(–50) cm, glabrous.

ascending or erect, rarely decumbent, (8–)16–50(–60) cm, glabrous, not glaucous.

Leaves

basal and cauline, glabrous;

basal and proximal cauline 20–80(–102) × 2–25 mm, blade spatulate to oblanceolate, base tapered, apex rounded to obtuse or acute, sometimes mucronate;

cauline 4–6 pairs, sessile, (16–)20–78 × 3–24 mm, blade ovate to elliptic or lanceolate, base clasping to cordate-clasping, apex acute to acuminate.

basal and cauline, or basal not persisting, ± leathery or not, glabrous, not glaucous;

basal and proximal cauline (18–)35–180 × (3–)6–15(–24) mm, blade oblanceolate to oblong or linear, base tapered, margins entire, apex obtuse to acute;

cauline 3–5(or 6) pairs, sessile or proximals sometimes short-petiolate, 34–115 × 2–14 mm, blade oblanceolate to oblong or linear, base tapered, apex obtuse to acute.

Thyrses

interrupted to nearly continuous, secund, 6–24(–31) cm, axis glabrous, verticillasters (2 or)3–10(–12), cymes 2–7-flowered;

proximal bracts ovate, 9–70 × 2–26 mm;

peduncles and pedicels glabrous.

interrupted or continuous, secund, (1–)6–18(–25) cm, axis glabrous or sparsely glandular-pubescent, verticillasters (2 or)3–7(–9), cymes (1 or)2–5-flowered, 2 per node;

proximal bracts lanceolate to linear, 25–70(–95) × (1–)3–12(–20) mm;

peduncles and pedicels glabrous or glandular-pubescent.

Flowers

calyx lobes ovate, 4–7 × 2.4–4 mm, margins erose, sometimes entire, broadly scarious, glabrous;

corolla blue to violet or lavender, rarely pink, with reddish or reddish purple nectar guides, tubular-funnelform to ventricose, 15–25 mm, glabrous externally, glabrous or sparsely to densely white-villous internally abaxially, tube 8–11 mm, throat gradually inflated, 4–7 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, 1–1.4 mm, sutures papillate;

staminode 10–13 mm, included or reaching orifice, 1–1.4 mm diam., tip abruptly recurved, bifurcate, distal 4–6 mm densely villous, hairs golden yellow, to 2 mm;

style 9–15 mm.

calyx lobes ovate to lanceolate, (4–)4.8–11 × 1.3–3.2(–3.7) mm, glabrous or glandular-pubescent;

corolla blue to lavender, with reddish purple nectar guides, ventricose, 18–33 mm, sparsely to moderately glandular-pubescent externally, glabrous, rarely sparsely white-villous, internally abaxially, tube 6–11 mm, throat gradually inflated, slightly constricted at orifice, 6–11 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, pollen sacs divergent to opposite, navicular to navicular-sigmoid, (1.3–)1.5–2.4 mm, dehiscing incompletely, proximal 1/5 indehiscent, connective not splitting, sides moderately pubescent, hairs white, to 0.8 mm, sutures denticulate, teeth to 0.1 mm;

staminode 11–19 mm, reaching orifice, 0.3–0.8 mm diam., tip straight to recurved, distal 3–7 mm sparsely to moderately villous, hairs yellow, to 0.8 mm;

style 12–22 mm.

Capsules

9–12 × 7–9 mm.

6–10 × 5–6.5 mm.

Penstemon secundiflorus

Penstemon scariosus

Distribution
from FNA
CO; NM; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

Penstemon scariosus comprises four varieties centered in the northwestern Colorado Plateau, Uinta Basin, and Wasatch Range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline leaf blades lanceolate to ovate; corollas light blue or lavender, rarely pink, sparsely to densely white-villous internally abaxially; Colorado, New Mexico, Wyoming.
var. secundiflorus
1. Cauline leaf blades oblanceolate to ovate or elliptic; corollas dark blue or violet, glabrous or sparsely white-villous internally abaxially; Colorado.
var. versicolor
1. Corollas 25–33 mm; calyx lobes 7–11 mm; pollen sacs 1.8–2.4 mm.
var. scariosus
1. Corollas 18–28 mm; calyx lobes (4–)4.8–8 mm; pollen sacs (1.3–)1.5–2.2 mm.
→ 2
2. Corollas sparsely to moderately glandular-pubescent externally, limbs blue to bluish purple; styles 12–14 mm.
var. cyanomontanus
2. Corollas sparsely glandular-pubescent externally, limbs lavender to pale blue or blue; styles 13–22 mm.
→ 3
3. Corolla limbs lavender to pale blue; styles 18–22 mm; basal leaves seldom persisting.
var. albifluvis
3. Corolla limbs blue; styles 13–17 mm; basal leaves usually persisting.
var. garrettii
Source FNA vol. 17, p. 124. FNA vol. 17, p. 178.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. secundiflorus var. secundiflorus, P. secundiflorus var. versicolor
P. scariosus var. albifluvis, P. scariosus var. cyanomontanus, P. scariosus var. garrettii, P. scariosus var. scariosus
Name authority Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 325. (1846) — (as Pentstemon) Pennell: Contr. U.S. Natl. Herb. 20: 353. (1920)
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