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side-bells beardtongue, sidebells penstemon

golden tongue penstemon, Malheur beardtongue, Malheur penstemon

Habit Plants: caudex sometimes herbaceous.
Stems

ascending to erect, (15–)20–45(–50) cm, glabrous.

ascending to erect, 8–28(–40) cm, retrorsely hairy.

Leaves

basal and cauline, glabrous;

basal and proximal cauline 20–80(–102) × 2–25 mm, blade spatulate to oblanceolate, base tapered, apex rounded to obtuse or acute, sometimes mucronate;

cauline 4–6 pairs, sessile, (16–)20–78 × 3–24 mm, blade ovate to elliptic or lanceolate, base clasping to cordate-clasping, apex acute to acuminate.

basal and cauline, not leathery, retrorsely hairy, hairs sometimes appressed, white, scalelike, sometimes basal and proximal cauline glabrate or glabrous abaxially;

basal and proximal cauline petiolate, 15–60(–78) × 4–12(–17) mm, blade spatulate to oblanceolate, base tapered, margins entire or dentate-serrate, apex rounded to obtuse or acute;

cauline 3 or 4 pairs, sessile or proximals short-petiolate, 20–50(–75) × 2–5(–10) mm, blade lanceolate to oblanceolate, base tapered to clasping, margins entire or obscurely and remotely dentate, apex obtuse to acute.

Thyrses

interrupted to nearly continuous, secund, 6–24(–31) cm, axis glabrous, verticillasters (2 or)3–10(–12), cymes 2–7-flowered;

proximal bracts ovate, 9–70 × 2–26 mm;

peduncles and pedicels glabrous.

± interrupted, cylindric, (3–)6–18 cm, axis glandular-pubescent and, sometimes, retrorsely hairy, verticillasters 3–6, cymes 2–5-flowered, 2 per node;

proximal bracts lanceolate, 18–46 × 2–12 mm;

peduncles and pedicels glandular-pubescent.

Flowers

calyx lobes ovate, 4–7 × 2.4–4 mm, margins erose, sometimes entire, broadly scarious, glabrous;

corolla blue to violet or lavender, rarely pink, with reddish or reddish purple nectar guides, tubular-funnelform to ventricose, 15–25 mm, glabrous externally, glabrous or sparsely to densely white-villous internally abaxially, tube 8–11 mm, throat gradually inflated, 4–7 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, 1–1.4 mm, sutures papillate;

staminode 10–13 mm, included or reaching orifice, 1–1.4 mm diam., tip abruptly recurved, bifurcate, distal 4–6 mm densely villous, hairs golden yellow, to 2 mm;

style 9–15 mm.

calyx lobes lanceolate, 5.5–8 × 1.2–2 mm, glandular-pubescent;

corolla light blue to bluish purple or magenta, with reddish violet nectar guides, ventricose, 14–22 mm, sparsely white-lanate internally abaxially, tube 5–7 mm, throat abruptly inflated, not constricted at orifice, (3–)5–7 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, pollen sacs opposite, explanate, 0.7–1(–1.2) mm, dehiscing completely, sutures smooth;

staminode 8–9 mm, exserted, 0.3–0.5 mm diam., tip straight to recurved, distal 5–7 mm moderately to densely pilose, hairs yellow or orangish yellow, to 1.2 mm;

style 9–10 mm.

Capsules

9–12 × 7–9 mm.

(6–)8–11 × 4.5–7 mm.

Penstemon secundiflorus

Penstemon miser

Phenology Flowering May–Jun.
Habitat Clayey soils, sagebrush shrublands, pine-juniper woodlands.
Elevation 800–1400 m. (2600–4600 ft.)
Distribution
from FNA
CO; NM; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; OR
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon miser is known from Owyhee County, Idaho, and southern Baker and northern Malheur counties, Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline leaf blades lanceolate to ovate; corollas light blue or lavender, rarely pink, sparsely to densely white-villous internally abaxially; Colorado, New Mexico, Wyoming.
var. secundiflorus
1. Cauline leaf blades oblanceolate to ovate or elliptic; corollas dark blue or violet, glabrous or sparsely white-villous internally abaxially; Colorado.
var. versicolor
Source FNA vol. 17, p. 124. FNA vol. 17, p. 140.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. secundiflorus var. secundiflorus, P. secundiflorus var. versicolor
Name authority Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 325. (1846) — (as Pentstemon) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. ed. 2, 2(1): 441. (1886) — (as Pentstemon)
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