side-bells beardtongue, sidebells penstemon
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Grinnell's beardtongue
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Plants: caudex sometimes herbaceous. |
Herbs. |
ascending to erect, (15–)20–45(–50) cm, glabrous. |
ascending to erect, 10–90 cm, glaucous or not. |
basal and cauline, glabrous; basal and proximal cauline 20–80(–102) × 2–25 mm, blade spatulate to oblanceolate, base tapered, apex rounded to obtuse or acute, sometimes mucronate; cauline 4–6 pairs, sessile, (16–)20–78 × 3–24 mm, blade ovate to elliptic or lanceolate, base clasping to cordate-clasping, apex acute to acuminate. |
basal and proximal cauline 31–90 × 11–20 mm, blade oblanceolate to elliptic, base tapered, margins finely to coarsely dentate, apex rounded to obtuse; cauline 3–6 pairs, short-petiolate or sessile, (15–)30–90 × 4–30 mm, blade lanceolate to ovate-oblong, base tapered, margins coarsely dentate, distals sometimes entire, apex obtuse or acute. |
interrupted to nearly continuous, secund, 6–24(–31) cm, axis glabrous, verticillasters (2 or)3–10(–12), cymes 2–7-flowered; proximal bracts ovate, 9–70 × 2–26 mm; peduncles and pedicels glabrous. |
interrupted, cylindric, (5–)7–34 cm, axis sparsely to densely glandular-pubescent, verticillasters 4–10, cymes 2–4-flowered; proximal bracts ovate to lanceolate, 5–50 × 2–18 mm; peduncles and pedicels spreading to ascending, glandular-pubescent. |
calyx lobes ovate, 4–7 × 2.4–4 mm, margins erose, sometimes entire, broadly scarious, glabrous; corolla blue to violet or lavender, rarely pink, with reddish or reddish purple nectar guides, tubular-funnelform to ventricose, 15–25 mm, glabrous externally, glabrous or sparsely to densely white-villous internally abaxially, tube 8–11 mm, throat gradually inflated, 4–7 mm diam., rounded abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, 1–1.4 mm, sutures papillate; staminode 10–13 mm, included or reaching orifice, 1–1.4 mm diam., tip abruptly recurved, bifurcate, distal 4–6 mm densely villous, hairs golden yellow, to 2 mm; style 9–15 mm. |
calyx lobes ovate, 4.5–8.5 × (2–)2.9–3.2 mm, glandular-pubescent; corolla white to pink or light lavender to light blue or light violet, with reddish purple nectar guides, strongly bilabiate, ventricose-ampliate, 22–35 mm, glandular-pubescent externally, glandular-pubescent internally, also white-villous abaxially, tube 5–8 mm, length 0.9–1.2 times calyx lobes, throat abruptly inflated, constricted at orifice, 10–18 mm diam., rounded abaxially; stamens included, filaments of shorter pair glandular-puberulent proximally, pollen sacs navicular to subexplanate, 1.6–2.2 mm, sutures smooth; staminode 20–22 mm, exserted, 0.4–0.8 mm diam., tip recurved, distal 8–10 mm densely lanate, hairs yellowish or whitish, to 4 mm; proximal 6–8 mm densely glandular-pubescent; style 15–18 mm, glabrous. |
9–12 × 7–9 mm. |
10–14 × 5–8 mm, glabrous. |
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= 16. |
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Flowering Apr–Jul. |
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Chaparral, foothills, pine-juniper woodlands, pine forests. |
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500–2700 m. (1600–8900 ft.) |
CO; NM; WY
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CA
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Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Two varieties of Penstemon grinnellii have been recognized. Variety grinnellii, with non-glaucous stems 10–60 cm and corollas white to pink and 22–30 mm, has been reported from the Western Transverse and Peninsular ranges, and in the San Bernardino, San Gabriel, and San Jacinto mountains. Variety scrophularioides, with glaucous stems 45–90 cm and corollas light lavender, light blue, or light violet and 26–35 mm, has been reported in the San Francisco Bay Area, southern Sierra Nevada and South Coast and Western Transverse ranges. Characteristics that distinguish the varieties overlap in some populations. Penstemon ×peirsonii Munz & I. M. Johnston, a putative hybrid between P. grinnellii and P. speciosus, and P. ×dubius Davidson, a putative hybrid between P. centranthifolius and P. grinnellii, both have been reported from California (D. D. Keck 1937b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Cauline leaf blades lanceolate to ovate; corollas light blue or lavender, rarely pink, sparsely to densely white-villous internally abaxially; Colorado, New Mexico, Wyoming. | var. secundiflorus |
1. Cauline leaf blades oblanceolate to ovate or elliptic; corollas dark blue or violet, glabrous or sparsely white-villous internally abaxially; Colorado. | var. versicolor |
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FNA vol. 17, p. 124. |
FNA vol. 17, p. 251. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. grinnellii subsp. scrophularioides, P. grinnellii var. scrophularioides, P. scrophularioides |
Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 325. (1846) — (as Pentstemon) |
Eastwood: Bull. Torrey Bot. Club 32: 207. (1905) — (as Pentstemon) |
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