side-bells beardtongue, sidebells penstemon
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foxglove beard-tongue, penstémon digitale, tall white beardtongue, talus slope penstemon
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Plants: caudex sometimes herbaceous. |
Herbs. |
ascending to erect, (15–)20–45(–50) cm, glabrous. |
erect, 25–90 cm, glabrous or sparsely retrorsely hairy, slightly glaucous or not. |
basal and cauline, glabrous; basal and proximal cauline 20–80(–102) × 2–25 mm, blade spatulate to oblanceolate, base tapered, apex rounded to obtuse or acute, sometimes mucronate; cauline 4–6 pairs, sessile, (16–)20–78 × 3–24 mm, blade ovate to elliptic or lanceolate, base clasping to cordate-clasping, apex acute to acuminate. |
basal and cauline, basal sometimes withering by anthesis, not leathery, glabrous; basal and proximal cauline 30–180(–250) × 4–38(–70) mm, blade spatulate to obovate or lanceolate, base tapered, margins entire or ± serrate or denticulate, apex rounded to obtuse or acute; cauline 5–8 pairs, petiolate or sessile, 26–195 × 4–55 mm, blade ovate to lanceolate, base tapered to clasping, margins entire or denticulate, apex acuminate to acute. |
interrupted to nearly continuous, secund, 6–24(–31) cm, axis glabrous, verticillasters (2 or)3–10(–12), cymes 2–7-flowered; proximal bracts ovate, 9–70 × 2–26 mm; peduncles and pedicels glabrous. |
interrupted, conic, 7–26(–34) cm, axis glabrous proximally, sparsely to densely glandular-pubescent distally, verticillasters (2 or)3–6, cymes (3–)5–12(–18)-flowered, 2 per node; proximal bracts lanceolate to linear, 9–105 × 1–40 mm, margins entire, sometimes serrulate; peduncles and pedicels spreading to ascending, sparsely glandular-pubescent. |
calyx lobes ovate, 4–7 × 2.4–4 mm, margins erose, sometimes entire, broadly scarious, glabrous; corolla blue to violet or lavender, rarely pink, with reddish or reddish purple nectar guides, tubular-funnelform to ventricose, 15–25 mm, glabrous externally, glabrous or sparsely to densely white-villous internally abaxially, tube 8–11 mm, throat gradually inflated, 4–7 mm diam., rounded abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, 1–1.4 mm, sutures papillate; staminode 10–13 mm, included or reaching orifice, 1–1.4 mm diam., tip abruptly recurved, bifurcate, distal 4–6 mm densely villous, hairs golden yellow, to 2 mm; style 9–15 mm. |
calyx lobes ovate, sometimes lanceolate, 4–8 × 2–3 mm, apex acute to acuminate, glandular-pubescent; corolla white, sometimes tinged lavender, with reddish purple nectar guides, ventricose, (17–)20–30 mm, glandular-pubescent externally, sparsely to moderately white-pubescent internally abaxially, tube 6–9 mm, throat abruptly inflated, 8–12 mm diam., slightly 2-ridged abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, navicular, 1.4–1.7 mm, dehiscing completely, connective splitting, sides sparsely pubescent, hairs white or purplish, to 0.6 mm, rarely glabrous, sutures papillate; staminode 13–17 mm, reaching orifice, 0.3–0.4 mm diam., tip straight to slightly recurved, distal 6–8 mm sparsely to moderately villous, hairs yellowish, to 1.5(–2) mm; style 13–18 mm. |
9–12 × 7–9 mm. |
8–14 × 4–6 mm, glabrous. |
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= 96. |
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Flowering Apr–Jul. |
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Prairies, meadows, roadsides, clearings in woods. |
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10–500 m. (0–1600 ft.) |
CO; NM; WY
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AL; AR; CT; DC; DE; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
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Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon digitalis appears to be native in the central Mississippi River Basin, with human activities expanding its range, particularly eastward (F. W. Pennell 1935). It is listed in VASCAN as introduced in the four Canadian provinces where it occurs (http://data.canadensys.net/vascan/taxon/7273). Plants resembling Penstemon digitalis but with smaller corollas (17–23 mm versus 23–30 mm), less glandular-pubescent inflorescences, less scarious-margined calyx lobes, and stems sometimes puberulent (versus glabrous) have been named P. alluviorum. Plants with those features come mostly from the southeastern Central Lowlands and north-central Coastal Plain provinces in the eastern United States in the south-central part of the range of P. digitalis (Alabama, Arkansas, Illinois, Indiana, Kentucky, Mississippi, Missouri, Ohio, and Tennessee). A. C. Koelling (1964) observed overlap in characters used to distinguish P. alluviorum from P. digitalis and concluded it was a small-flowered variant of P. digitalis. Specimens of Penstemon digitalis with lanceolate and acuminate-tipped calyx lobes can be mistaken for P. calycosus, but calyx lobe length and, usually, the presence of hairs on the pollen sacs allow them to be accurately identified. Penstemon digitalis is widely cultivated and spreads readily. R. R. Clinebell and P. Bernhardt (1998) found evidence that plants are self-compatible. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Cauline leaf blades lanceolate to ovate; corollas light blue or lavender, rarely pink, sparsely to densely white-villous internally abaxially; Colorado, New Mexico, Wyoming. | var. secundiflorus |
1. Cauline leaf blades oblanceolate to ovate or elliptic; corollas dark blue or violet, glabrous or sparsely white-villous internally abaxially; Colorado. | var. versicolor |
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FNA vol. 17, p. 124. |
FNA vol. 17, p. 202. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. alluviorum |
Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 325. (1846) — (as Pentstemon) |
Nuttall ex Sims: Bot. Mag. 52: plate 2587. (1825) — (as Pentstemon) |
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