Penstemon sect. Elmigera |
Penstemon labrosus |
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San Gabriel beardtongue |
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Habit | Herbs. | |||||||||||||
Stems | glabrous or retrorsely hairy, glaucous or not. |
erect, sometimes ascending, 30–80 cm, glabrous, slightly glaucous. |
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Leaves | basal and cauline, opposite, leathery or not, glabrous, puberulent, or retrorsely hairy, glaucous or not; basal and proximal cauline petiolate; cauline sessile or short-petiolate, blade subcordate, ovate, spatulate, oblanceolate, lanceolate, elliptic, or linear, margins entire. |
glabrous, slightly glaucous; basal and proximal cauline 35–110 × 3–11 mm, blade oblanceolate to lanceolate, base tapered, margins entire, apex obtuse to acute; cauline 3–6 pairs, sessile or proximals short-petiolate, 27–85 × 1–6 mm, blade linear, base tapered, apex acute, rarely obtuse. |
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Thyrses | interrupted, ± secund to secund, rarely cylindric, axis glabrous, sometimes retrorsely hairy, cymes 2 per node, sometimes 1 per node (P. labrosus); peduncles and pedicels spreading to ascending or erect. |
± secund, 13–30 cm, axis glabrous, verticillasters 5–10, cymes 1- or 2(or 3)-flowered; proximal bracts linear, 5–30 × 1–2 mm; peduncles and pedicels ascending to erect, glabrous. |
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Flowers | calyx lobes: margins entire or erose, broadly scarious, sometimes scarious (P. labrosus), glabrous or glandular-pubescent; corolla red to scarlet or orange, rarely yellow, bilaterally symmetric or nearly radially symmetric (P. eatonii), weakly to strongly bilabiate, not personate, salverform, tubular-funnelform, or tubular, glabrous externally, glabrous or hairy internally abaxially, throat slightly to gradually inflated, not constricted to constricted at orifice, rounded to slightly 2-ridged abaxially; stamens included to exserted, filaments glabrous, pollen sacs parallel, divergent, or opposite, navicular, dehiscing incompletely, proximal 1/4–1/2 indehiscent, connective not splitting, sides glabrous or hispidulous to lanate-villous; sutures papillate or denticulate, papillae or teeth to 0.1 mm; staminode included or slightly exserted, flattened distally, 0.3–0.9 mm diam., tip straight, rarely recurved, glabrous or distal 5–20% hairy, hairs to 1.5 mm; style glabrous. |
calyx lobes ovate to lanceolate, 3.5–6 × 1.8–2.4 mm, margins erose, glabrous; corolla red to scarlet or orange, rarely yellow, without nectar guides, bilaterally symmetric, strongly bilabiate, salverform, 30–40 mm, glabrous internally, tube 5–7 mm, throat 4–6 mm diam., not constricted at orifice, rounded abaxially, abaxial lobes spreading to reflexed, adaxial lobes projecting, galeate; stamens exserted (but concealed by projecting adaxial lip), pollen sacs opposite, proximal 1/4–1/3 indehiscent, 1.6–2 mm, sides glabrous, sutures papillate; staminode 11–14 mm, included, tip straight, glabrous; style 20–28 mm, exserted. |
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Capsules | glabrous. |
7–10 × 4–6 mm. |
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Seeds | brown to dark brown, angled, 1.4–4.2 mm. |
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2n | = 16. |
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Penstemon sect. Elmigera |
Penstemon labrosus |
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Phenology | Flowering Jun–Sep. | |||||||||||||
Habitat | Pinyon-juniper woodlands, pine and mixed-hardwood forests. | |||||||||||||
Elevation | 1200–2400 m. (3900–7900 ft.) | |||||||||||||
Distribution | w United States; Mexico |
CA; Mexico (Baja California)
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Discussion | Species 8 (4 in the flora). Members of sect. Elmigera all exhibit the syndrome of characteristics usually associated with hummingbird pollination. Section Elmigera is not monophyletic; species in the flora area probably are related to species in sect. Glabri (C. A. Wessinger et al. 2016). Four species usually included in sect. Elmigera, but not in the flora area, are Penstemon henricksonii Straw, P. imberbis (Kunth) Trautvetter, P. luteus G. L. Nesom, and P. wislizeni (A. Gray) Straw; all occur in Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon labrosus is known from the Peninsular and Transverse ranges of southern California and mountain ranges of Baja California. In California, populations have been documented in Inyo, Kern, Los Angeles, Riverside, San Bernardino, San Diego, and Ventura counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 145. | FNA vol. 17, p. 149. | ||||||||||||
Parent taxa | ||||||||||||||
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Synonyms | Elmigera | P. barbatus var. labrosus | ||||||||||||
Name authority | (Reichenbach ex Spach) Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 329. (1846) — (as Pentstemon) | (A. Gray) Hooker f.: Bot. Mag. 110: plate 6738. (1884) | ||||||||||||
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