Penstemon sect. Elmigera |
Penstemon cardinalis |
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cardinal beardtongue, scarlet beardtongue |
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Habit | Herbs. | |||||||||||||||||
Stems | glabrous or retrorsely hairy, glaucous or not. |
ascending to erect, 32–100 cm, glabrous, slightly glaucous. |
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Leaves | basal and cauline, opposite, leathery or not, glabrous, puberulent, or retrorsely hairy, glaucous or not; basal and proximal cauline petiolate; cauline sessile or short-petiolate, blade subcordate, ovate, spatulate, oblanceolate, lanceolate, elliptic, or linear, margins entire. |
glabrous, not glaucous; basal and proximal cauline 40–80(–150) × 14–40(–60) mm, blade spatulate to obovate, oblanceolate, or elliptic, base tapered, margins entire, apex rounded to obtuse; cauline 3–7 pairs, sessile, (11–)20–135 × 9–50 mm, blade spatulate to elliptic, ovate, or subcordate, base truncate to broadly cordate-clasping, apex rounded to acute or mucronate. |
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Thyrses | interrupted, ± secund to secund, rarely cylindric, axis glabrous, sometimes retrorsely hairy, cymes 2 per node, sometimes 1 per node (P. labrosus); peduncles and pedicels spreading to ascending or erect. |
secund, (3–)6–44 cm, axis glabrous, verticillasters 3–15, cymes 2–4(–8)-flowered; proximal bracts suborbiculate, ovate, or lanceolate, (4–)16–60 × (2–)5–45 mm; peduncles and pedicels erect, glabrous. |
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Flowers | calyx lobes: margins entire or erose, broadly scarious, sometimes scarious (P. labrosus), glabrous or glandular-pubescent; corolla red to scarlet or orange, rarely yellow, bilaterally symmetric or nearly radially symmetric (P. eatonii), weakly to strongly bilabiate, not personate, salverform, tubular-funnelform, or tubular, glabrous externally, glabrous or hairy internally abaxially, throat slightly to gradually inflated, not constricted to constricted at orifice, rounded to slightly 2-ridged abaxially; stamens included to exserted, filaments glabrous, pollen sacs parallel, divergent, or opposite, navicular, dehiscing incompletely, proximal 1/4–1/2 indehiscent, connective not splitting, sides glabrous or hispidulous to lanate-villous; sutures papillate or denticulate, papillae or teeth to 0.1 mm; staminode included or slightly exserted, flattened distally, 0.3–0.9 mm diam., tip straight, rarely recurved, glabrous or distal 5–20% hairy, hairs to 1.5 mm; style glabrous. |
calyx lobes ovate, 1.6–6 × 1.4–3 mm, margins entire or erose, glabrous; corolla red to crimson, with reddish violet nectar guides, bilaterally symmetric, weakly bilabiate, tubular, 22–30 mm, densely yellow-villous internally abaxially, tube 6–8 mm, throat 6–8 mm diam., constricted at orifice, rounded abaxially, abaxial lobes spreading to reflexed, adaxial lobes projecting to erect; stamens included, pollen sacs divergent, 1.5–1.9 mm, sides glabrous, sutures denticulate, teeth to 0.1 mm; staminode 15–17 mm, included or slightly exserted, tip straight to recurved, distal 1–3 mm sparsely lanate, hairs yellow, to 1.5 mm; style 18–28 mm, usually exserted. |
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Capsules | glabrous. |
10–15 × 5–9 mm. |
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Seeds | brown to dark brown, angled, 1.4–4.2 mm. |
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Penstemon sect. Elmigera |
Penstemon cardinalis |
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Distribution | w United States; Mexico |
NM; TX
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Discussion | Species 8 (4 in the flora). Members of sect. Elmigera all exhibit the syndrome of characteristics usually associated with hummingbird pollination. Section Elmigera is not monophyletic; species in the flora area probably are related to species in sect. Glabri (C. A. Wessinger et al. 2016). Four species usually included in sect. Elmigera, but not in the flora area, are Penstemon henricksonii Straw, P. imberbis (Kunth) Trautvetter, P. luteus G. L. Nesom, and P. wislizeni (A. Gray) Straw; all occur in Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Penstemon cardinalis occurs as two weakly differentiated varieties in isolated ranges of the Southern Rocky Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 145. | FNA vol. 17, p. 147. | ||||||||||||||||
Parent taxa | ||||||||||||||||||
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Synonyms | Elmigera | |||||||||||||||||
Name authority | (Reichenbach ex Spach) Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 329. (1846) — (as Pentstemon) | Wooton & Standley: Contr. U.S. Natl. Herb. 16: 171. (1913) — (as Pentstemon) | ||||||||||||||||
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