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lance-leaf beardtongue

Habit Herbs.
Stems

retrorsely hairy or puberulent, sometimes glandular-pubescent distally or glabrescent, not glaucous.

24–75(–98) cm, sparsely to densely retrorsely hairy or puberulent, usually also glandular-pubescent distally.

Leaves

basal and cauline, basal usually absent at anthesis, opposite, leathery or not, glabrous, puberulent, or retrorsely hairy, not glaucous;

basal and proximal cauline petiolate, sometimes short-petiolate or sessile (P. dasyphyllus);

cauline sessile, sometimes short-petiolate (P. lanceolatus), blade lanceolate or linear, margins entire.

retrorsely hairy;

basal and proximal cauline 8–80 × 3–12 mm, blade lanceolate or spatulate, base tapered, margins entire, apex rounded to obtuse or acute;

cauline 7–15 pairs, 9–85(–110) × 1–11(–22) mm, blade lanceolate to linear, base tapered to truncate, apex acute to acuminate, rarely obtuse.

Thyrses

interrupted, ± secund to cylindric, axis glandular-pubescent or glabrous, cymes 1 or 2 per node;

peduncles and pedicels ascending to erect.

± secund, 5–36 cm, axis glandular-pubescent, verticillasters 3–6, cymes 1 or 2(or 3)-flowered;

proximal bracts lanceolate to linear, 5–16 × 1–3 mm;

peduncles and pedicels glandular-pubescent, sometimes also retrorsely hairy.

Flowers

calyx lobes: margins entire or erose (P. stenophyllus), herbaceous or narrowly scarious, sometimes broadly scarious (P. stenophyllus), glandular-pubescent or glabrous;

corolla lavender to blue, violet, purple, red, or scarlet, bilaterally symmetric, bilabiate, not personate, ventricose or tubular-funnelform, glandular-pubescent externally, glabrous or glandular-pubescent internally abaxially, throat abruptly or gradually inflated, not constricted at orifice, rounded to 2-ridged abaxially;

stamens included to exserted, filaments glabrous, pollen sacs parallel to divergent, navicular to slightly saccate, dehiscing completely or incompletely, distal 1/5 or less sometimes indehiscent, connective splitting, sides glabrous, sutures denticulate, teeth to 0.2–0.3 mm;

staminode included or exserted, flattened distally, 0.3–2.6 mm diam., tip recurved, glabrous;

style glabrous.

calyx lobes ovate to lanceolate, 2.9–7 × 1.5–2.5(–3.6) mm, glandular-pubescent;

corolla scarlet to red, unlined internally or lined on abaxial surface with faint reddish purple nectar guides, tubular-funnelform, 22–35 mm, glandular-pubescent internally abaxially, tube 5–6 mm, throat gradually inflated, 6–8 mm diam., rounded abaxially;

stamens exserted (hidden by galeate adaxial lobes), pollen sacs parallel to divergent, 1.5–2 mm, sutures denticulate, teeth to 0.2 mm;

staminode 15–17 mm, included, 0.3–0.4 mm diam.;

style 25–30 mm, usually barely exserted from galea.

Capsules

glabrous.

9–14 × 6–7 mm.

Seeds

brown to dark brown, angled, 1.9–3.2 mm.

2n

= 16.

Penstemon sect. Chamaeleon

Penstemon lanceolatus

Phenology Flowering May–Sep.
Habitat Gravelly pinyon-juniper woodlands, pine woodlands, thorn scrub, desert grasslands.
Elevation 1200–1800 m. (3900–5900 ft.)
Distribution
sw United States; Mexico
from FNA
AZ; NM; TX; Mexico (Aguascalientes, Chihuahua, Coahuila, Durango, Nayarit, Nuevo León, San Luis Potosí, Tamaulipas, Veracruz, Zacatecas)
Discussion

Species 4 (3 in the flora).

Crosswhite circumscribed sect. Chamaeleon to include a group of southern species distinguished by incompletely dehiscent pollen sacs with prominently denticulate sutures. Penstemon punctatus Brandegee, a species not treated by Crosswhite, might belong here; the only specimens of that species seen were from Coahuila, Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon lanceolatus is known from southern Arizona (Cochise, Graham, and Greenlee counties), southern New Mexico (Doña Ana, Grant, Hidalgo, Luna, and Sierra counties), and western Texas (Brewster County).

Crosswhite published Penstemon ramosus as an avowed substitute for the homonym P. pauciflorus Greene, distinguishing P. ramosus from P. lanceolatus by the former’s branched stems and relatively narrower leaves (1 mm wide versus 4–8 mm wide) with revolute margins. He mapped P. ramosus in southeastern Arizona and southwestern New Mexico and P. lanceolatus in western Texas and northern Mexico. Morphologic differences between these taxa are not consistent enough to warrant recognition of P. ramosus (J. L. Anderson et al. 2007).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas scarlet to red, tubular-funnelform, glandular-pubescent internally abaxially.
P. lanceolatus
1. Corollas violet to blue, lavender, or purple, ventricose, glabrous internally.
→ 2
2. Thyrse axes glandular-pubescent.
P. dasyphyllus
2. Thyrse axes glabrous.
P. stenophyllus
Source FNA vol. 17, p. 108. FNA vol. 17, p. 109.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon Plantaginaceae > Penstemon > subg. Penstemon > sect. Chamaeleon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. dasyphyllus, P. lanceolatus, P. stenophyllus
Synonyms P. ramosus
Name authority Crosswhite: Sida 2: 339. (1966) Bentham: Pl. Hartw., 22. (1839) — (as Pentstemon)
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