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cataiina beardtongue, Catalina beardtongue
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Subshrubs, usually cespitose. |
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retrorsely hairy, rarely glabrous, not glaucous. |
ascending to erect, 10–35 cm, glabrous or sparsely retrorsely hairy, hairs white, scalelike. |
cauline, rarely basal and cauline (P. retrorsus), opposite, leathery or not, glabrous, glabrate, pubescent, scabrous, puberulent, or retrorsely hairy, not glaucous; basal and proximal cauline petiolate; cauline petiolate or sessile, blade obovate, spatulate, oblanceolate, lanceolate, elliptic, or linear, margins entire. |
leathery, glabrous, rarely sparsely retrorsely hairy, hairs white, scalelike abaxially, densely puberulent, hairs white, scalelike adaxially; cauline 10–30 pairs, petiolate, 5–18 × 1–1.5 mm, blade oblanceolate to linear, base tapered, apex mucronate. |
continuous or interrupted, cylindric or secund, axis retrorsely hairy, glandular-pubescent, or puberulent and glandular-pubescent, rarely glabrous, cymes 1 or 2 per node; peduncles and pedicels spreading to ascending or erect. |
continuous or interrupted, ± secund, 2–15 cm, axis glandular-pubescent, verticillasters 5–13, cymes 1–3-flowered, 1 per node; proximal bracts oblanceolate to linear, 5–12 × 0.5–1 mm; peduncles and pedicels ascending to erect, glandular-pubescent. |
calyx tube 0.2–1 mm (1.5–2 mm in P. retrorsus), lobes: margins entire or erose, ± scarious or herbaceous, glandular-pubescent or retrorsely hairy, rarely glabrate; corolla lavender, blue, violet, or purple, rarely scarlet, bilaterally symmetric, bilabiate, not personate, funnelform, tubular-funnelform, salverform, ventricose-ampliate, or ampliate, glandular-pubescent externally, hairy internally abaxially, rarely glabrous, throat slightly to abruptly inflated, not constricted at orifice, 2-ridged abaxially; stamens included to exserted, filaments glabrous, rarely sparsely glandular-puberulent proximally, pollen sacs opposite, rarely divergent, navicular to subexplanate or explanate, dehiscing completely, connective splitting, sides glabrous, sutures papillate or smooth; staminode included to exserted, ± flattened distally, 0.1–0.5 mm diam., tip straight to recurved, distal (20–)50–100% hairy, hairs to 0.7–1.3 mm; style glabrous. |
calyx lobes ovate to lanceolate, 2.5–3.5 × 1.2–1.8 mm, scarious-margined, glandular-pubescent, hairs pointed; corolla lavender to violet or purple, lined internally abaxially with reddish purple nectar guides, tubular-funnelform, 9–14 mm, densely yellow- or whitish lanate internally abaxially, tube 3–5 mm, throat gradually inflated, 3.5–5 mm diam., slightly 2-ridged abaxially; stamens included, filaments glabrous, pollen sacs opposite, navicular, 0.5–0.8 mm, sutures papillate; staminode 7–8 mm, included or reaching orifice, flattened distally, 0.2–0.4 mm diam., tip straight to recurved, distal 5–6 mm densely pilose, hairs golden yellow, to 1 mm; style 8–11 mm. |
glabrous. |
5–8 × 3–4.8 mm. |
dark brown to black, angled to rounded, 1.1–2.1 mm. |
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Flowering Jun–Aug. |
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Pinyon-juniper and oak woodlands, granite crevices. |
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1900–2100 m. (6200–6900 ft.) |
w United States; nw Mexico |
AZ |
Species 11 (11 in the flora). Most species of sect. Caespitosi formed a clade in the molecular study by A. D. Wolfe et al. (2006); placement of the clade within subg. Penstemon is not clear. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon discolor is known from the Atascosa, Dragoon, Santa Catalina, and Santa Teresa mountains in Cochise, Graham, Pima, and Santa Cruz counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Corollas salverform, scarlet (throats sometimes yellow or yellow- or orange-spotted). | P. pinifolius |
1. Corollas tubular-funnelform, funnelform, ampliate, or ventricose-ampliate, lavender, blue, violet, or purple. | → 2 |
2. Leaves glabrous or retrorsely hairy, hairs appressed, white, scalelike. | → 3 |
3. Stems prostrate, spreading, or ascending; cauline leaves 1–5 pairs. | → 4 |
4. Leaves densely retrorsely hairy, hairs appressed, white, scalelike; pollen sacs 0.8–1.2(–1.4) mm; Arizona, California, Nevada, Utah. | P. thompsoniae |
4. Leaves glabrous or sparsely hairy, hairs appressed, white, scalelike abaxially, ± densely hairy, hairs appressed, white, scalelike adaxially; pollen sacs 0.6–0.8 mm; Utah. | P. xylus |
3. Stems ascending to erect; cauline leaves 6–30 pairs. | → 5 |
5. Leaves glabrous, rarely sparsely retrorsely hairy, hairs white, scalelike abaxially, densely puberulent, hairs white, scalelike adaxially; corollas 9–14 mm; pollen sacs 0.5–0.8 mm; Arizona. | P. discolor |
5. Leaves sparsely to densely retrorsely hairy, hairs appressed, white, scalelike; corollas 14–20 mm; pollen sacs 0.9–1.4 mm; Arizona, California, New Mexico. | → 6 |
6. Corollas tubular-funnelform, 14–18 mm, purple to violet; California. | P. californicus |
6. Corollas ventricose-ampliate, 16–20 mm, blue to lavender; Arizona, New Mexico. | P. linarioides |
2. Leaves glabrous, glabrate, or hairy, hairs pointed. | → 7 |
7. Calyx tubes 1.5–2 mm; proximal bracts oblanceolate. | P. retrorsus |
7. Calyx tubes 0.2–1 mm; proximal bracts oblanceolate to linear. | → 8 |
8. Calyx lobes herbaceous- or narrowly scarious-margined. | → 9 |
9. Stems and leaves not cinereous; leaf blades obovate to spatulate, oblanceolate, or linear; Arizona, Colorado, Utah, Wyoming. | P. caespitosus |
9. Stems and leaves cinereous; leaf blades linear; Colorado. | P. teucrioides |
8. Calyx lobes broadly scarious-margined. | → 10 |
10. Corollas ventricose-ampliate; calyx lobe apices acute to acuminate. | P. linarioides |
10. Corollas funnelform; calyx lobe apices acuminate to caudate. | → 11 |
11. Leaf blades linear, rarely oblanceolate; stems ascending to erect; corollas glabrous or sparsely white-villous internally abaxially; Utah. | P. abietinus |
11. Leaf blades elliptic to obovate, oblanceolate, or linear; stems prostrate, decumbent, ascending, or erect; corollas sparsely to densely yellow-lanate or yellow-pilose internally abaxially; Colorado, New Mexico, Utah. | P. crandallii |
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FNA vol. 17, p. 98. |
FNA vol. 17, p. 103. |
Plantaginaceae > Penstemon > subg. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi |
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P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. caespitosus, P. californicus, P. crandallii, P. discolor, P. linarioides, P. pinifolius, P. retrorsus, P. teucrioides, P. thompsoniae, P. xylus |
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P. |
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(Rydberg) Pennell: Contr. U.S. Natl. Herb. 20: 327, 334. (1920) |
D. D. Keck: Bull. Torrey Bot. Club 64: 379. (1937) |
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