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Garrett's beardtongue, White River beardtongue

long-sepal beardtongue

Habit Herbs.
Stems

ascending or erect, rarely decumbent, (8–)16–50(–60) cm, glabrous, not glaucous.

erect, (40–)60–120 cm, puberulent, at least proximally, slightly glaucous or not.

Leaves

basal and cauline, or basal not persisting, ± leathery or not, glabrous, not glaucous;

basal and proximal cauline (18–)35–180 × (3–)6–15(–24) mm, blade oblanceolate to oblong or linear, base tapered, margins entire, apex obtuse to acute;

cauline 3–5(or 6) pairs, sessile or proximals sometimes short-petiolate, 34–115 × 2–14 mm, blade oblanceolate to oblong or linear, base tapered, apex obtuse to acute.

basal and cauline, basal often withering by anthesis, not leathery, glabrous;

basal and proximal cauline 47–110 × 10–24 mm, blade oblanceolate, base tapered, margins entire or ± serrate, apex rounded to obtuse;

cauline 3–7 pairs, sessile or proximals short-petiolate, (14–)44–155 × (2–)13–46 mm, blade lanceolate to ovate, proximals sometimes oblanceolate, base clasping or tapered, margins serrate, apex acute to acuminate.

Thyrses

interrupted or continuous, secund, (1–)6–18(–25) cm, axis glabrous or sparsely glandular-pubescent, verticillasters (2 or)3–7(–9), cymes (1 or)2–5-flowered, 2 per node;

proximal bracts lanceolate to linear, 25–70(–95) × (1–)3–12(–20) mm;

peduncles and pedicels glabrous or glandular-pubescent.

interrupted, conic, (5–)14–20 cm, axis sparsely puberulent and glandular-pubescent, rarely glabrous, verticillasters 2–5, cymes (1–)5–15-flowered, 2 per node;

proximal bracts ovate to lanceolate, sometimes linear, 13–80 × (2–)5–32 mm, margins serrate, sometimes entire;

peduncles and pedicels spreading to ascending, sparsely glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, (4–)4.8–11 × 1.3–3.2(–3.7) mm, glabrous or glandular-pubescent;

corolla blue to lavender, with reddish purple nectar guides, ventricose, 18–33 mm, sparsely to moderately glandular-pubescent externally, glabrous, rarely sparsely white-villous, internally abaxially, tube 6–11 mm, throat gradually inflated, slightly constricted at orifice, 6–11 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, pollen sacs divergent to opposite, navicular to navicular-sigmoid, (1.3–)1.5–2.4 mm, dehiscing incompletely, proximal 1/5 indehiscent, connective not splitting, sides moderately pubescent, hairs white, to 0.8 mm, sutures denticulate, teeth to 0.1 mm;

staminode 11–19 mm, reaching orifice, 0.3–0.8 mm diam., tip straight to recurved, distal 3–7 mm sparsely to moderately villous, hairs yellow, to 0.8 mm;

style 12–22 mm.

calyx lobes lanceolate, 5–9 × 0.8–1.9 mm, apex acuminate, sparsely glandular-pubescent;

corolla pale lavender to violet, with faint violet nectar guides, ventricose, 20–35 mm, glandular-pubescent externally, ± white-pubescent internally abaxially, tube 5–7 mm, throat abruptly inflated, 8–11 mm diam., slightly 2-ridged abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, navicular, 1.2–1.4 mm, dehiscing completely, connective splitting, sides glabrous, rarely sparsely pubescent, hairs white, to 0.2 mm, sutures papillate;

staminode 11–13 mm, reaching orifice, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 2–5 mm sparsely pubescent, hairs yellowish, to 1 mm;

style 14–16 mm.

Capsules

6–10 × 5–6.5 mm.

7–8 × 4–5 mm, glabrous.

2n

= 96.

Penstemon scariosus

Penstemon calycosus

Phenology Flowering May–Jun.
Habitat Woodlands, meadows, rocky slopes, stream banks.
Elevation 10–300 m. (0–1000 ft.)
Distribution
from FNA
CO; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DC; GA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NJ; NY; OH; PA; RI; SC; TN; VA; VT; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Penstemon scariosus comprises four varieties centered in the northwestern Colorado Plateau, Uinta Basin, and Wasatch Range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon calycosus is concentrated in the Central Lowlands and Interior Low Plateaus; occurrences farther south and east, especially in New England, may be from introductions (F. W. Pennell 1935). The range lies almost entirely within that of P. digitalis, but historic ranges of both P. calycosus and P. digitalis are difficult to delimit due to both past and continuing introduction and spread of both species. Calyx lobe lengths to 12 mm are reported in the literature; those appear to be from fruiting calyces. Penstemon calycosus usually has glabrous anthers, but many specimens, especially from Illinois and Ohio, bear hairs on the adaxial surfaces of the anthers. Some populations from this region exhibit variation in vegetative (leaf size and bract shape and margins) and floral characters (calyx lobe shape and size, corolla color, and anther pubescence), suggesting possible hybridization with P. digitalis, an observation that would be consistent with the findings of A. C. Koelling (1964).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas 25–33 mm; calyx lobes 7–11 mm; pollen sacs 1.8–2.4 mm.
var. scariosus
1. Corollas 18–28 mm; calyx lobes (4–)4.8–8 mm; pollen sacs (1.3–)1.5–2.2 mm.
→ 2
2. Corollas sparsely to moderately glandular-pubescent externally, limbs blue to bluish purple; styles 12–14 mm.
var. cyanomontanus
2. Corollas sparsely glandular-pubescent externally, limbs lavender to pale blue or blue; styles 13–22 mm.
→ 3
3. Corolla limbs lavender to pale blue; styles 18–22 mm; basal leaves seldom persisting.
var. albifluvis
3. Corolla limbs blue; styles 13–17 mm; basal leaves usually persisting.
var. garrettii
Source FNA vol. 17, p. 178. FNA vol. 17, p. 197.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. scariosus var. albifluvis, P. scariosus var. cyanomontanus, P. scariosus var. garrettii, P. scariosus var. scariosus
Name authority Pennell: Contr. U.S. Natl. Herb. 20: 353. (1920) Small: Bull. Torrey Bot. Club 25: 470. (1898) — (as Pentstemon)
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