cliff beardtongue, cliff penstemon, rock penstemon
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toad-flax beardtongue, toadflax penstemon
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Subshrubs, cespitose. |
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ascending to erect, 3–16 cm, puberulent to pubescent, not glaucous. |
ascending to erect, 8–50 cm, retrorsely hairy, hairs pointed or white, scalelike. |
persistent, 3–8 pairs, distals usually distinctly smaller than proximals, short-petiolate or sessile, 4–22 × 3–14 mm, blade round to elliptic or spatulate, base tapered to cuneate, margins ± serrate, apex rounded to obtuse, rarely acute, glabrous or hairy, usually glaucous. |
not leathery, glabrous, glabrate, or sparsely to densely retrorsely hairy, hairs pointed, sometimes appressed, white, scalelike; cauline 6–12(–20) pairs, petiolate or sessile, 4–26 × 0.5–3.5 mm, blade oblanceolate to lanceolate or linear, base tapered to truncate, apex mucronate. |
continuous, ± secund, 1–3 cm, axis hairy proximally, pubescent or glandular-pubescent distally, verticillasters 1–3, cymes 1-flowered; proximal bracts elliptic to ovate, 3–9 × 2–6 mm; peduncles and pedicels ascending to erect, glandular-pubescent. |
continuous or interrupted, ± secund, 3–17 cm, axis glandular-pubescent, verticillasters 3–9(–12), cymes 1 or 2(or 3)-flowered, 1(or 2) per node; proximal bracts lanceolate to linear, 6–25 × 0.7–1 mm; peduncles and pedicels ascending to erect, retrorsely hairy, hairs pointed, and sparsely glandular-pubescent. |
calyx lobes lanceolate-elliptic to oblong, 6–11 × 2–2.8 mm, glandular-pubescent; corolla pink to red, pinkish lavender, or rose red, essentially unlined internally but abaxial ridges usually white, not personate, funnelform, 25–36 mm, glabrous externally, glabrous internally or sparsely white-lanate abaxially, tube 8–12 mm, throat 6–8 mm diam.; stamens: longer pair exserted, pollen sacs 1–1.5 mm; staminode 13–15 mm, slightly flattened distally, 0.1 mm diam., tip straight, glabrous or distal 1–2 mm sparsely villous, hairs yellow, to 0.8 mm; style 26–30 mm. |
calyx lobes ovate, rarely lanceolate, 4–7(–9) × 2–3 mm, broadly scarious-margined, apex acute to acuminate, glandular-pubescent, hairs pointed; corolla blue or lavender, lined internally abaxially with reddish purple nectar guides, ventricose-ampliate, 16–20 mm, moderately white- or yellowish villous internally abaxially, tube 5–6 mm, throat abruptly inflated, 5.5–8 mm diam., slightly 2-ridged abaxially; stamens: longer pair reaching orifice or slightly exserted, filaments glabrous, pollen sacs opposite, navicular, 0.9–1.3 mm, sutures papillate; staminode 6–8 mm, exserted, flattened distally, 0.3–0.5 mm diam., tip straight to recurved, distal 1–5 mm densely pilose, hairs yellow or golden yellow, to 1.2 mm, rest of distal 1–7 mm glabrous or sparsely to moderately pilose; style 10–11 mm. |
7–9 × 5–7 mm. |
5–9 × 3.5–5 mm. |
= 16. |
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Flowering Jun–Jul. |
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Cliffs, rock outcrops. |
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60–2400 m. (200–7900 ft.) |
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CA; OR; WA
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AZ; CO; NM; NV; UT
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Penstemon rupicola occurs largely in the Siskiyou Mountains and Cascade Range from northern California to northern Washington. It is partly sympatric with and forms hybrids with P. cardwellii, P. davidsonii var. davidsonii, and P. fruticosus var. fruticosus (A. D. Every 1977). Hybrids between P. rupicola and P. davidsonii var. davidsonii are encountered frequently in the vicinity of Crater Lake National Park and Mt. Hood (Every). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Penstemon linarioides is widespread and highly variable. D. D. Keck (1937) recognized seven subspecies on the basis of habit, pubescence, leaf shape, and staminode bearding. Some of the variation on which those subspecies are based appears to be clinal or too variable within or among regional populations to be of taxonomic value. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Leaves glabrous, glabrate, retrorsely hairy, or pubescent, hairs pointed. | var. sileri |
1. Leaves sparsely to densely retrorsely hairy, hairs appressed, white, scalelike. | → 2 |
2. Cauline leaf blades oblanceolate, distals sometimes lanceolate, 1.5–3.5 mm wide. | var. maguirei |
2. Cauline leaf blades oblanceolate to linear, 0.5–2 mm wide. | → 3 |
3. Staminodes: distal 3–5 mm densely pilose, hairs golden yellow or yellow, to 1 mm, rest of distal 1–4 mm sparsely to moderately pilose. | var. linarioides |
3. Staminodes: distal 1 mm densely pilose, hairs yellow, to 0.8 mm, rest of distal 3–4 mm glabrous or sparsely pilose (with much shorter hairs). | var. coloradoensis |
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FNA vol. 17, p. 92. |
FNA vol. 17, p. 103. |
Plantaginaceae > Penstemon > subg. Dasanthera > sect. Erianthera |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. newberryi var. rupicola |
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(Piper) Howell: Fl. N.W. Amer., 510. (1901) — (as Pentstemon) |
A. Gray: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 112. (1859) — (as Pentstemon) |
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