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Wassuk beardtongue, Wassuk Range beardtongue

superb beardtongue

Habit Herbs.
Stems

erect, 50–120 cm, glaucous.

ascending to erect, 30–140(–160) cm, glaucous.

Leaves

basal and proximal cauline 25–100 × (5–)10–35 mm, blade oblanceolate, base tapered, margins coarsely dentate, apex obtuse to acute;

cauline 3–5 pairs, sessile, 25–65 × 12–25 mm, blade lanceolate, base sessile, clasping or distal ones connate-perfoliate, margins coarsely dentate, apex acute.

glabrous, glaucous;

basal and proximal cauline 40–160 × 14–40 mm, blade spatulate to oblanceolate or elliptic, base tapered, margins entire, apex rounded to obtuse or acute;

cauline 3–8 pairs, sessile, (17–)44–115 × (4–)23–45 mm, blade ovate to oblanceolate or lanceolate, base tapered to cordate-clasping to connate-perfoliate, margins entire, apex rounded or obtuse to acute.

Thyrses

interrupted, secund, 20–60 cm, axis glabrous, sometimes sparsely glandular-pubescent distally, verticillasters 10–16, cymes 2- or 3-flowered;

proximal bracts lanceolate, 15–60 × 4–22 mm;

peduncles and pedicels ascending to erect, glandular-pubescent.

interrupted, cylindric, (15–)30–60(–90) cm, axis glabrous, verticillasters 9–12(–20), cymes (1–)3–9-flowered;

proximal bracts ovate to lanceolate, 11–55 × 4–30 mm;

peduncles and pedicels ascending to erect, glabrous or pedicels sparsely glandular-pubescent.

Flowers

calyx lobes ovate, 4–5.5 × 2–2.7 mm, glandular-pubescent;

corolla pink to rose pink, with reddish nectar guides, strongly bilabiate, ventricose to ventricose-ampliate, (27–)30–35 mm, glandular-pubescent externally, sparsely white-lanate internally abaxially, not glandular-pubescent, tube 7–10 mm, length 1.8–2 times calyx lobes, throat gradually inflated, not constricted at orifice, 10–16 mm diam., rounded abaxially;

stamens included or longer pair exserted, filaments glabrous, pollen sacs navicular, 1.2–1.5 mm, sutures smooth;

staminode 22–24 mm, exserted, 0.4–0.6 mm diam., tip recurved, distal 6–7 mm pilose, hairs yellow or golden yellow, to 2 mm;

style 18–24 mm, glabrous.

calyx lobes ovate to lanceolate, 2.8–5 × 1.5–2.1 mm, margins entire or erose, sparsely glandular-pubescent proximally, sometimes glabrous;

corolla orangish pink to red, without nectar guides, nearly radially symmetric, weakly bilabiate, tubular-funnelform, 17–22 mm, glandular-pubescent externally, glandular-pubescent internally abaxially, tube 5–7 mm, throat slightly inflated, 4–6 mm diam., rounded abaxially;

stamens included, pollen sacs explanate, 1–1.5 mm, sutures smooth;

staminode 8–11 mm, flattened distally, 0.6–1 mm diam., tip straight, distal 2–3 mm retrorsely hairy, hairs yellow or whitish, to 1.5 mm;

style 10–11 mm.

Capsules

7–11 × 4–6 mm, glabrous.

10–13 × 5–8 mm.

2n

= 16.

Penstemon rubicundus

Penstemon superbus

Phenology Flowering May–Sep. Flowering (Mar–)Apr–Jun.
Habitat Rocky or gravelly slopes, drainages, roadsides, sagebrush or shadscale shrublands, pine-juniper woodlands. Gravelly or rocky canyons, slopes, washes, desert grasslands, pinyon-juniper and oak woodlands.
Elevation 1300–2100 m. (4300–6900 ft.) 900–1800 m. (3000–5900 ft.)
Distribution
from FNA
NV
[BONAP county map]
from FNA
AZ; NM; Mexico (Chihuahua, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon rubicundus is known from the Pine Nut Mountains (Douglas County) and the Wassuk Range (Mineral County) of western Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon superbus is known from Cochise, Gila, Graham, Greenlee, and Pima counties, Arizona, Grant and Hidalgo counties, New Mexico, and in adjacent Chihuahua and Sonora, Mexico. The species resembles P. parryi; it differs by broader leaves and corollas that are orangish pink to red, more nearly radially symmetric, and without white hairs abaxially in the throats. Penstemon superbus also resembles P. alamosensis, which is known from New Mexico and Texas to the east of P. superbus. The glabrous staminode of P. alamosensis distinguishes it from P. superbus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 254. FNA vol. 17, p. 154.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Synonyms P. puniceus
Name authority D. D. Keck: Amer. Midl. Naturalist 18: 802. (1937) A. Nelson: Proc. Biol. Soc. Wash. 17: 100. (1904) — (as Pentstemon)
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