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Wassuk beardtongue, Wassuk Range beardtongue

Absaroka Range beardtongue

Habit Herbs.
Stems

erect, 50–120 cm, glaucous.

ascending to erect, 7–27 cm, glabrous or ± puberulent distally or at nodes, not glaucous.

Leaves

basal and proximal cauline 25–100 × (5–)10–35 mm, blade oblanceolate, base tapered, margins coarsely dentate, apex obtuse to acute;

cauline 3–5 pairs, sessile, 25–65 × 12–25 mm, blade lanceolate, base sessile, clasping or distal ones connate-perfoliate, margins coarsely dentate, apex acute.

essentially cauline, basal leaves absent or reduced, ± leathery, glabrous, not glaucous;

basal and proximal cauline 12–90(–105) × 3–23 mm, blade elliptic-lanceolate to oblanceolate, base tapered, margins entire, apex obtuse to acute;

cauline 2–4 pairs, petiolate or sessile, 35–95(–107) × 11–30 mm, blade elliptic-lanceolate to oblanceolate, base tapered to truncate, apex obtuse to acute.

Thyrses

interrupted, secund, 20–60 cm, axis glabrous, sometimes sparsely glandular-pubescent distally, verticillasters 10–16, cymes 2- or 3-flowered;

proximal bracts lanceolate, 15–60 × 4–22 mm;

peduncles and pedicels ascending to erect, glandular-pubescent.

continuous, ± secund, 3–15 cm, axis glabrous or ± glandular, verticillasters 1–7, cymes 2–8-flowered, 2 per node;

proximal bracts ovate to elliptic, 16–85 × 7–31 mm;

peduncles and pedicels glabrous or ± glandular, peduncles to 15 mm, pedicels 1–3 mm.

Flowers

calyx lobes ovate, 4–5.5 × 2–2.7 mm, glandular-pubescent;

corolla pink to rose pink, with reddish nectar guides, strongly bilabiate, ventricose to ventricose-ampliate, (27–)30–35 mm, glandular-pubescent externally, sparsely white-lanate internally abaxially, not glandular-pubescent, tube 7–10 mm, length 1.8–2 times calyx lobes, throat gradually inflated, not constricted at orifice, 10–16 mm diam., rounded abaxially;

stamens included or longer pair exserted, filaments glabrous, pollen sacs navicular, 1.2–1.5 mm, sutures smooth;

staminode 22–24 mm, exserted, 0.4–0.6 mm diam., tip recurved, distal 6–7 mm pilose, hairs yellow or golden yellow, to 2 mm;

style 18–24 mm, glabrous.

calyx lobes ovate to lanceolate, 6.5–12 × 2.8–4.5 mm, margins erose, apex acuminate to caudate, ± glandular;

corolla blue to violet, with violet nectar guides, ventricose, 18–33 mm, glabrous externally, glabrous internally, tube 7–9 mm, throat abruptly inflated, slightly constricted at orifice, 7–9 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, pollen sacs divergent to nearly opposite, navicular-sigmoid, (1.8–)2.2–2.6 mm, dehiscing incompletely, connective not splitting, sides sparsely to moderately hispidulous or pubescent, hairs white, to 0.3 mm;

sutures denticulate, teeth to 0.1 mm;

staminode 13–17 mm, included, 0.5–0.9 mm diam., tip straight, glabrous or distal 1–3 mm pubescent, hairs yellow, to 0.5 mm;

style 17–20 mm.

Capsules

7–11 × 4–6 mm, glabrous.

8–12 × 5–8 mm.

Penstemon rubicundus

Penstemon absarokensis

Phenology Flowering May–Sep. Flowering Jun–Jul.
Habitat Rocky or gravelly slopes, drainages, roadsides, sagebrush or shadscale shrublands, pine-juniper woodlands. Volcanic talus slopes.
Elevation 1300–2100 m. (4300–6900 ft.) 1800–3000 m. (5900–9800 ft.)
Distribution
from FNA
NV
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Penstemon rubicundus is known from the Pine Nut Mountains (Douglas County) and the Wassuk Range (Mineral County) of western Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon absarokensis is known from the Absaroka Range of northwestern Wyoming (Fremont and Park counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 254. FNA vol. 17, p. 161.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority D. D. Keck: Amer. Midl. Naturalist 18: 802. (1937) Evert: Madroño 31: 140, fig. 1. (1984)
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