cut-leaf beardtongue, Richardson's beardtongue, Richardson's penstemon, shrubby penstemon
|
Sunset Crater beardtongue, Sunset Crater penstemon
|
Subshrubs. |
Herbs. |
ascending to erect, 20–95 cm, retrorsely hairy, hairs pointed, not glaucous. |
ascending to erect, 30–60 cm, glaucous. |
cauline, opposite or subopposite, glabrate or retrorsely hairy, not glaucous; cauline 5–13 pairs, short-petiolate or sessile, 13–70 × 5–30 mm, blade ovate to lanceolate, base tapered, margins laciniate-dentate to irregularly laciniate-pinnatifid, apex obtuse to acute. |
basal and proximal cauline 26–56 × 7–15 mm, blade lanceolate to elliptic, base tapered, margins coarsely dentate or serrate, apex obtuse to acute; cauline 4–7 pairs, short-petiolate or sessile, 30–55 × 5–32 mm, blade ovate to lanceolate, base tapered or auriculate-clasping to connate-perfoliate on distal leaves, margins coarsely dentate, apex acute to acuminate. |
interrupted, cylindric, 2–25(–48) cm, axis densely glandular-pubescent, verticillasters 3–7(–15), cymes 2–7-flowered, 1 or 2 per node; proximal bracts lanceolate, 7–43 × 2–16 mm; peduncles and pedicels spreading, sparsely to densely glandular-pubescent. |
interrupted, secund, 8–30 cm, axis glabrous or sparsely glandular-pubescent distally, verticillasters 3–10, cymes 1–3-flowered; proximal bracts ovate to triangular, 12–35 × 8–32 mm; peduncles and pedicels ascending to erect, glandular-pubescent. |
calyx lobes ovate to lanceolate, 4–7 × 1.3–2.3 mm, glandular-pubescent; corolla bluish to lavender, or pinkish, with reddish purple nectar guides, ventricose-ampliate, 15–32 mm, glandular-pubescent externally, glabrous internally or sparsely to moderately white-lanate abaxially, tube 7–12 mm, throat gradually inflated, 5–14 mm diam., rounded abaxially; stamens: longer pair reaching orifice or exserted, filaments glabrous, pollen sacs parallel, 1.3–2 mm, distal 1/2 indehiscent, sides glabrous, sutures denticulate, teeth to 0.2 mm; staminode 16–23 mm, exserted, 0.3–0.8 mm diam., glabrous or distal 2–8 mm sparsely lanate, hairs yellow, to 2 mm; style 11–23 mm. |
calyx lobes ovate to elliptic, 3.5–5.5 × 2.5–3.8 mm, glandular-pubescent; corolla pink to rose pink, with reddish purple nectar guides, strongly bilabiate, ventricose, 23–27 mm, glandular-pubescent externally, glandular-pubescent internally, also usually moderately white-lanate abaxially, tube 6–10 mm, length 1.6–2 times calyx lobes, throat abruptly inflated, ± constricted at orifice, 9–12 mm diam., rounded abaxially; stamens: longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs explanate, 1.3–1.6 mm, sutures smooth; staminode 16–18 mm, exserted, 0.4–0.6(–1) mm diam., tip straight, glabrous or distal 2–4 mm moderately lanate, hairs yellow, to 1 mm; style 17–22 mm, glabrous. |
5–9 × 4–5 mm. |
9–12 × 6–7 mm, glabrous. |
= 16. |
= 16. |
|
|
|
|
|
Flowering Apr–Aug. |
|
Volcanic cinder fields, pine forests. |
|
1900–2600 m. (6200–8500 ft.) |
OR; WA; BC
|
AZ
|
Varieties 3 (3 in the flora). Penstemon richardsonii consists of three weakly differentiated varieties that are partially sympatric on the eastern slope of the Cascade Range. The Okanagan-Coville tribe of northeastern Washington and the Paiute tribe of the Great Basin region used Penstemon richardsonii as a drug plant to treat typhoid fever and as a dermatological aid (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon clutei occurs only near Indian Flat and Sunset Crater in south-central Coconino County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Corollas 15–20 mm; styles 11–15 mm; staminodes glabrous or sparsely hairy distally. | var. curtiflorus |
1. Corollas 22–32 mm; styles 16–23 mm; staminodes with distal 2–8 mm sparsely lanate, rarely glabrous. | → 2 |
2. Leaf blades ovate to lanceolate (in outline), margins laciniate-dentate to irregularly laciniate-pinnatifid. | var. richardsonii |
2. Leaf blades ovate, sometimes lanceolate, margins dentate, rarely laciniate-dentate. | var. dentatus |
|
|
FNA vol. 17, p. 241. |
FNA vol. 17, p. 249. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
|
|
|
P. pseudospectabilis subsp. clutei |
Douglas ex Lindley: Bot. Reg. 13: plate 1121. (1828) — (as Pentstemon) |
A. Nelson: Amer. Bot. (Binghamton) 33: 109. (1927) |
| |