mat-root beardtongue, matroot penstemon
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derived penstemon, Plumas County beardtongue
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Herbs. |
Subshrubs. |
ascending to erect, (15–)20–42 cm, retrorsely hairy or puberulent, not glaucous. |
ascending to erect, 20–80 cm, glabrous, sometimes ± puberulent, hairs pointed, glaucous. |
essentially cauline, basal absent or poorly developed, not leathery, ± puberulent; cauline (4 or)5–8 pairs, sessile, (20–)30–65 × (2–)4–10(–20) mm, blade ovate to lanceolate, base tapered, margins entire, apex obtuse to acute. |
basal and cauline, basal sometimes few, opposite, glabrous, sometimes proximals puberulent, hairs pointed, glaucous; basal and proximal cauline 15–80 × 2–10 mm, blade spatulate to oblanceolate, base tapered, margins entire, apex rounded to obtuse or acute; cauline 4–6 pairs, sessile, 20–80 × 3–9 mm, blade ovate to lanceolate, base tapered, margins entire, apex acute. |
interrupted or continuous, cylindric, 5–14 cm, axis glandular-pubescent, verticillasters (2 or)3–6, cymes 2–4(–8)-flowered, 2 per node; proximal bracts lanceolate to linear, 7–46 × 1–10(–20) mm, margins entire; peduncles and pedicels ascending to erect, glandular-pubescent. |
interrupted, cylindric, 5–25 cm, axis glandular-pubescent, verticillasters 4–6, cymes 1–3-flowered, 2 per node; proximal bracts ovate to lanceolate, 4–50 × 1–8 mm; peduncles and pedicels ascending, glandular-pubescent. |
calyx lobes lanceolate, 5–8 × 1.2–2.5 mm, glandular-pubescent; corolla blue to dark blue, purple, or violet, with reddish purple nectar guides, bilabiate, not personate, funnelform, 16–23 mm, glandular-pubescent externally, moderately yellowish lanate internally abaxially, tube 4.5–6 mm, throat slightly inflated, 4–6 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular to subexplanate, 0.8–1.3 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 10–12 mm, included, 0.3–0.4 mm diam., tip straight, distal 1 mm densely pilose or lanate, hairs yellow, to 1.4 mm, remaining distal 4–5 mm sparsely pilose, hairs yellow, to 0.5 mm; style 11–14 mm. |
calyx lobes ovate to lanceolate, 3.8–6 × 1.5–2.2 mm, glandular-pubescent; corolla violet to blue or purple, without nectar guides, ventricose to ventricose-ampliate, 25–38 mm, glandular-pubescent externally, glabrous internally, tube 7–9 mm, throat gradually inflated, 8–10 mm diam., 2-ridged abaxially; stamens included, filaments glabrous, pollen sacs parallel, 2.4–3.2 mm, distal 1/5–1/3 indehiscent, sides pilose to lanate, hairs white, to 0.6 mm near filament attachment, sutures denticulate, teeth to 0.2 mm; staminode 11–15 mm, included, 0.4–0.6 mm diam., glabrous; style 15–20 mm. |
5–8 × 3.5–5 mm, glabrous. |
7–9 × 4–5 mm. |
= 16. |
= 64. |
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Flowering May–Jul. |
Flowering May–Aug(–Oct). |
Rocky flats, slopes, ravines, sagebrush shrublands, pine woodlands. |
Volcanic soils, scrub, open pine and fir forests. |
1500–2400 m. (4900–7900 ft.) |
1000–2200 m. (3300–7200 ft.) |
CO; ID; MT; NV; UT; WY
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CA
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Penstemon radicosus is known from the Central Rocky Mountains and mountain ranges in the northern Great Basin. The species is most likely to be confused with P. laxus or P. watsonii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon neotericus is known from the Cascade Range and northern Sierra Nevada of northern California (Butte, Lassen, Placer, Plumas, Shasta, Sierra, Siskiyou, Tehama, and Yuba counties). D. D. Keck (1932) used morphologic and geographic data to hypothesize that P. neotericus, an octoploid, arose through hybridization between P. azureus and P. laetus. J. Clausen (1933) provided cytological evidence supporting that hypothesis. A diploid chromosome number of 2n = 32 listed by N. H. Holmgren (1993) is probably a transcription error. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 219. |
FNA vol. 17, p. 238. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
A. Nelson: Bull. Torrey Bot. Club 25: 280. (1898) — (as Pentstemon) |
D. D. Keck: Univ. Calif. Publ. Bot. 16: 398, fig. 13. (1932) |
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