mat-root beardtongue, matroot penstemon
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California beardtongue, California penstemon
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Herbs. |
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ascending to erect, (15–)20–42 cm, retrorsely hairy or puberulent, not glaucous. |
ascending to erect, (5–)8–30 cm, retrorsely hairy, hairs white, scalelike. |
essentially cauline, basal absent or poorly developed, not leathery, ± puberulent; cauline (4 or)5–8 pairs, sessile, (20–)30–65 × (2–)4–10(–20) mm, blade ovate to lanceolate, base tapered, margins entire, apex obtuse to acute. |
± leathery or not, densely retrorsely hairy, hairs appressed, white, scalelike; cauline 10–30 pairs, petiolate, 8–16 × 1.5–2.5 mm, blade oblanceolate, base tapered, apex mucronate. |
interrupted or continuous, cylindric, 5–14 cm, axis glandular-pubescent, verticillasters (2 or)3–6, cymes 2–4(–8)-flowered, 2 per node; proximal bracts lanceolate to linear, 7–46 × 1–10(–20) mm, margins entire; peduncles and pedicels ascending to erect, glandular-pubescent. |
interrupted or continuous, secund, (1–)2–9 cm, axis retrorsely hairy with appressed, white, scalelike hairs, verticillasters 2–13, cymes 1–4-flowered, 1 per node; proximal bracts oblanceolate, 2–10 × 0.5–1.5 mm; peduncles and pedicels spreading to ascending, retrorsely hairy with appressed, white, scalelike hairs, and sparsely glandular-pubescent. |
calyx lobes lanceolate, 5–8 × 1.2–2.5 mm, glandular-pubescent; corolla blue to dark blue, purple, or violet, with reddish purple nectar guides, bilabiate, not personate, funnelform, 16–23 mm, glandular-pubescent externally, moderately yellowish lanate internally abaxially, tube 4.5–6 mm, throat slightly inflated, 4–6 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular to subexplanate, 0.8–1.3 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 10–12 mm, included, 0.3–0.4 mm diam., tip straight, distal 1 mm densely pilose or lanate, hairs yellow, to 1.4 mm, remaining distal 4–5 mm sparsely pilose, hairs yellow, to 0.5 mm; style 11–14 mm. |
calyx lobes ovate, 3.5–5.5 × 1.5–2 mm, scarious-margined, retrorsely hairy with appressed, white, scalelike hairs, and sparsely glandular-pubescent; corolla purple to violet, lined internally abaxially with reddish purple nectar guides, tubular-funnelform, 14–18 mm, sparsely white-lanate internally abaxially, tube 5–6 mm, throat gradually inflated, 4.5–6 mm diam., 2-ridged abaxially; stamens included, filaments glabrous or sparsely puberulent and sometimes with appressed, white, scalelike hairs proximally, pollen sacs opposite, navicular, 0.9–1.4 mm, sutures papillate; staminode 7–9 mm, reaching orifice or exserted, flattened distally, 0.3–0.4 mm diam., tip straight to recurved, distal 4–5 mm pilose, hairs yellow, to 0.7 mm; style 9–11 mm. |
5–8 × 3.5–5 mm, glabrous. |
5–7 × 4–5 mm. |
= 16. |
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Flowering May–Jul. |
Flowering May–Jul(–Aug). |
Rocky flats, slopes, ravines, sagebrush shrublands, pine woodlands. |
Sandy or gravelly soils, pine-juniper woodlands, pine forests. |
1500–2400 m. (4900–7900 ft.) |
1200–2300 m. (3900–7500 ft.) |
CO; ID; MT; NV; UT; WY
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CA; Mexico (Baja California)
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Penstemon radicosus is known from the Central Rocky Mountains and mountain ranges in the northern Great Basin. The species is most likely to be confused with P. laxus or P. watsonii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In California, Penstemon californicus is known from the San Jacinto Mountains in Orange and Riverside counties; it also occurs in the Sierra San Pedro Mártir in Baja California. The chromosome number of 2n = 16 reported by N. H. Holmgren (1993) is likely an error based on a count for Scrophularia californica reported in P. H. Raven et al. (1965) and later misattributed to P. californicus by D. V. Clark (1971) as n = 48 (the count listed in Raven et al.), and then corrected to 2n = 16. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 219. |
FNA vol. 17, p. 101. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. linarioides var. californicus |
A. Nelson: Bull. Torrey Bot. Club 25: 280. (1898) — (as Pentstemon) |
(Munz & I. M. Johnston) D. D. Keck: Bull. Torrey Bot. Club 64: 378. (1937) |
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