Penstemon pseudospectabilis |
Penstemon sect. Spectabiles |
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desert penstemon, Mojave beardtongue |
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Habit | Herbs. | Herbs, shrubs, or subshrubs. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | ascending to erect, (30–)60–100 cm, slightly glaucous or not. |
glabrous, glaucous, sometimes not. |
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Leaves | basal and proximal cauline 20–165 × 12–50 mm, blade ovate to lanceolate, base tapered, margins finely to coarsely dentate, apex obtuse to acute; cauline 4–7 pairs, sessile or proximals short-petiolate, 45–90 × 22–76 mm, blade ovate to triangular, base tapered or sessile on proximals, auriculate-clasping to connate-perfoliate on distal ones, margins finely to coarsely dentate, apex acute. |
basal and cauline, basals sometimes few (P. clevelandii, P. stephensii) or absent (P. clevelandii), or leaves essentially cauline (P. fruticiformis, P. incertus), opposite, leathery, sometimes not (P. pseudospectabilis, P. spectabilis, P. stephensii), glabrous, glaucous, sometimes not (P. clevelandii, P. spectabilis, P. stephensii); basal and proximal cauline petiolate; cauline sessile or short-petiolate, blade cordate, ovate, triangular-ovate, triangular, oblanceolate, triangular-lanceolate, lanceolate, ovate-oblong, or linear, margins entire or toothed. |
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Thyrses | interrupted or continuous, secund, (6–)12–48 cm, axis glabrous, verticillasters 4–13, cymes 1–3(–6)-flowered; proximal bracts ovate (connate-perfoliate) or lanceolate (auriculate-clasping), (9–)16–55 × 10–60 mm; peduncles and pedicels ascending to erect, glabrous or glandular-pubescent. |
interrupted, rarely continuous, cylindric or secund, axis glabrous or glandular-pubescent, cymes 2 per node; peduncles and pedicels spreading to ascending or erect. |
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Flowers | calyx lobes ovate, 3–5.5 × 2–3.2 mm, glabrous or glandular-pubescent; corolla rose pink, with reddish purple nectar guides, strongly bilabiate, ventricose, (18–)25–34 mm, glandular-pubescent externally, glandular-pubescent internally, tube 6–9 mm, length 1.6–2 times calyx lobes, throat gradually inflated, ± constricted at orifice, 7–10 mm diam., rounded abaxially; stamens: longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs explanate, 1.5–2 mm, sutures smooth; staminode 11–17 mm, included, 0.2–0.4 mm diam., tip straight to recurved, glabrous or distal 1–2 mm sparsely to moderately lanate, hairs yellow, to 1 mm; style 20–25 mm, glabrous. |
calyx lobes: margins entire or erose, sometimes denticulate (P. spectabilis), ± scarious, sometimes herbaceous (P. pseudospectabilis), glabrous or glandular-pubescent; corolla white to pink, rose, red, magenta, yellow, lavender, blue, violet, or purple, bilaterally symmetric, bilabiate to strongly bilabiate, not personate, ventricose or ventricose-ampliate, sometimes tubular-funnelform or funnelform, glandular-pubescent externally, sometimes glabrous, glabrous or hairy internally abaxially, glandular-pubescent on both surfaces, or glandular-pubescent on both surfaces and hairy abaxially, throat gradually to abruptly inflated, constricted or not at orifice, rounded abaxially, sometimes 2-ridged; stamens included or longer pair reaching orifice or exserted, filaments glabrous or shorter pair glandular-puberulent proximally, pollen sacs opposite, rarely divergent (P. incertus), navicular to subexplanate or explanate, dehiscing completely, rarely incompletely (P. incertus), connective splitting, rarely not (P. incertus), sides glabrous, sutures smooth, rarely papillate or denticulate, teeth to 0.1 mm; staminode included or exserted, flattened distally, 0.2–1.1 mm diam., tip straight to recurved, rarely coiled, glabrous or distal 10–50% hairy, hairs to 4 mm, sometimes proximal 10–40% hairy, hairs glandular; style glabrous, rarely sparsely glandular-pubescent proximally. |
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Capsules | 7–12 × 4–6 mm, glabrous. |
glabrous or sparsely glandular-pubescent distally. |
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Seeds | brown to dark brown or black, angled, rarely ± rounded, 1–4 mm. |
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Penstemon pseudospectabilis |
Penstemon sect. Spectabiles |
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Distribution |
AZ; CA; NM; UT; n Mexico
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w United States; n Mexico |
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Discussion | Varieties 2 (2 in the flora). Two varieties, distinguished on the basis of inflorescence indument, are largely allopatric. Penstemon ×crideri A. Nelson, a putative hybrid between P. eatonii and P. pseudospectabilis, has been reported from Arizona (A. Nelson 1936). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 16 (12 in the flora). D. D. Keck (1937b) circumscribed sect. Peltanthera, a name that is illegitimate and superfluous, to include species from southwestern United States and northern Mexico with showy, white to pink, red, or purple, tubular to ventricose corollas and completely dehiscent, glabrous pollen sacs. His section encompasses species included here mostly in sect. Spectabiles and species treated here in sects. Gentianoides and Petiolati. Core species of sect. Spectabiles, those with ventricose corollas and treated by Keck in so-called subsect. Spectabiles (not validly published), were monophyletic in a molecular study (C. A. Wessinger et al. 2016) that included nine of the 12 species treated here in sect. Spectabiles. That study also provided support for inclusion of Penstemon alamosensis and P. parryi, species with tubular corollas and treated here in sect. Gentianoides, in sect. Spectabiles. However, broader taxon sampling is needed before definitive circumscription of sects. Gentianoides and Spectabiles is possible. Four species and one variety in sect. Spectabiles occur in Mexico: P. angelicus (I. M. Johnston) Moran, P. eximius D. D. Keck, P. rotundifolius A. Gray, P. spectabilis var. subinteger (D. D. Keck) C. C. Freeman, and P. vizcainensis Moran. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 253. | FNA vol. 17, p. 245. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles | Plantaginaceae > Penstemon > subg. Penstemon | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | M. E. Jones: Contr. W. Bot. 12: 66. (1908) — (as Pentstemon) | Pennell: Contr. U.S. Natl. Herb. 20: 326, 329. (1920) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |