desert penstemon, Mojave beardtongue
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pine-leaf beardtongue, pineneedle beardtongue, pineneedle penstemon
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Herbs. |
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ascending to erect, (30–)60–100 cm, slightly glaucous or not. |
erect, (5–)10–50 cm, glabrous or retrorsely hairy, hairs pointed. |
basal and proximal cauline 20–165 × 12–50 mm, blade ovate to lanceolate, base tapered, margins finely to coarsely dentate, apex obtuse to acute; cauline 4–7 pairs, sessile or proximals short-petiolate, 45–90 × 22–76 mm, blade ovate to triangular, base tapered or sessile on proximals, auriculate-clasping to connate-perfoliate on distal ones, margins finely to coarsely dentate, apex acute. |
leathery, sparsely retrorsely hairy, hairs pointed, especially along margins, or glabrous; cauline 10–30 pairs, sessile, 4–32 × 0.5–1.2 mm, blade linear, base tapered to clasping, apex acuminate. |
interrupted or continuous, secund, (6–)12–48 cm, axis glabrous, verticillasters 4–13, cymes 1–3(–6)-flowered; proximal bracts ovate (connate-perfoliate) or lanceolate (auriculate-clasping), (9–)16–55 × 10–60 mm; peduncles and pedicels ascending to erect, glabrous or glandular-pubescent. |
interrupted, secund, (1–)3–15 cm, axis retrorsely hairy to puberulent, or glabrous, verticillasters 1–7, cymes 1–3-flowered, 1(or 2) per node; proximal bracts linear, 3–14 × 0.3–1 mm; peduncles and pedicels ascending to erect, glandular-pubescent, sometimes puberulent. |
calyx lobes ovate, 3–5.5 × 2–3.2 mm, glabrous or glandular-pubescent; corolla rose pink, with reddish purple nectar guides, strongly bilabiate, ventricose, (18–)25–34 mm, glandular-pubescent externally, glandular-pubescent internally, tube 6–9 mm, length 1.6–2 times calyx lobes, throat gradually inflated, ± constricted at orifice, 7–10 mm diam., rounded abaxially; stamens: longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs explanate, 1.5–2 mm, sutures smooth; staminode 11–17 mm, included, 0.2–0.4 mm diam., tip straight to recurved, glabrous or distal 1–2 mm sparsely to moderately lanate, hairs yellow, to 1 mm; style 20–25 mm, glabrous. |
calyx lobes lanceolate, 5–7 × 1.5–2 mm, scarious-margined, glandular-pubescent, hairs pointed; corolla scarlet, throat sometimes yellow or yellow- or orange-spotted, unlined internally, salverform, 25–32 mm, yellow-lanate internally abaxially, tube 4–6 mm, throat slightly inflated, 4–5 mm diam., slightly 2-ridged abaxially; stamens exserted (often hidden by galeate adaxial lobes), filaments glabrous, pollen sacs opposite, explanate, 0.8–1 mm, sutures smooth; staminode 12–17 mm, included, slightly flattened distally, 0.1–0.2 mm diam., tip straight, distal 9–10 mm densely pilose, hairs yellow, to 1.3 mm; style 21–27 mm. |
7–12 × 4–6 mm, glabrous. |
5–9 × 3–4 mm. |
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= 16. |
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Flowering Jun–Jul(–Sep). |
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Rocky slopes, cliffs. |
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1800–3100 m. (5900–10200 ft.) |
AZ; CA; NM; UT; n Mexico
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AZ; NM; Mexico (Chihuahua, Sonora)
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Varieties 2 (2 in the flora). Two varieties, distinguished on the basis of inflorescence indument, are largely allopatric. Penstemon ×crideri A. Nelson, a putative hybrid between P. eatonii and P. pseudospectabilis, has been reported from Arizona (A. Nelson 1936). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon pinifolius has been included as an anomalous member of sect. Leptostemon Trautvetter; R. M. Straw (1962) expressed doubt about that association. It was allied with members of sect. Caespitosi in the molecular study by A. D. Wolfe et al. (2006). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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FNA vol. 17, p. 253. |
FNA vol. 17, p. 105. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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M. E. Jones: Contr. W. Bot. 12: 66. (1908) — (as Pentstemon) |
Greene: Bot. Gaz. 6: 218. (1881) — (as Pentstemon) |
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