desert penstemon, Mojave beardtongue
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Canaan Mountain beardtongue, sandloving penstemon
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Herbs. |
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ascending to erect, (30–)60–100 cm, slightly glaucous or not. |
decumbent to ascending, 5–26 cm, usually ± fistulose, glutinous and covered with sand, not glaucous. |
basal and proximal cauline 20–165 × 12–50 mm, blade ovate to lanceolate, base tapered, margins finely to coarsely dentate, apex obtuse to acute; cauline 4–7 pairs, sessile or proximals short-petiolate, 45–90 × 22–76 mm, blade ovate to triangular, base tapered or sessile on proximals, auriculate-clasping to connate-perfoliate on distal ones, margins finely to coarsely dentate, apex acute. |
basal and cauline, not leathery, glutinous and covered with sand, not glaucous; basal and proximal cauline 18–80 × 4–15 mm, blade oblanceolate, base tapered, margins entire, crisped, apex obtuse to acute; cauline 2–5 pairs, sessile, 15–75 × 2–8 mm, blade oblanceolate to lanceolate, base tapered to truncate, margins crisped, apex rounded or obtuse to acute. |
interrupted or continuous, secund, (6–)12–48 cm, axis glabrous, verticillasters 4–13, cymes 1–3(–6)-flowered; proximal bracts ovate (connate-perfoliate) or lanceolate (auriculate-clasping), (9–)16–55 × 10–60 mm; peduncles and pedicels ascending to erect, glabrous or glandular-pubescent. |
continuous, cylindric, 4–15 cm, axis glutinous and covered with sand, verticillasters 4–10, cymes 1–6-flowered, 2 per node; proximal bracts lanceolate, 14–45 × 2–5(–8) mm; peduncles and pedicels glutinous and covered with sand. |
calyx lobes ovate, 3–5.5 × 2–3.2 mm, glabrous or glandular-pubescent; corolla rose pink, with reddish purple nectar guides, strongly bilabiate, ventricose, (18–)25–34 mm, glandular-pubescent externally, glandular-pubescent internally, tube 6–9 mm, length 1.6–2 times calyx lobes, throat gradually inflated, ± constricted at orifice, 7–10 mm diam., rounded abaxially; stamens: longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs explanate, 1.5–2 mm, sutures smooth; staminode 11–17 mm, included, 0.2–0.4 mm diam., tip straight to recurved, glabrous or distal 1–2 mm sparsely to moderately lanate, hairs yellow, to 1 mm; style 20–25 mm, glabrous. |
calyx lobes lanceolate, 5–6.5 × 1–2 mm, glutinous and covered with sand; corolla lavender to blue-lavender, with violet nectar guides, weakly ventricose, 14–17 mm, glabrous externally, usually glutinous and covered with sand distally, especially on lobes, glabrous internally, tube 5–6 mm, throat gradually inflated, not constricted at orifice, 5–6 mm diam., slightly 2-ridged abaxially; stamens: longer pair reaching orifice, pollen sacs opposite, navicular, 0.9–1.4 mm, dehiscing completely or incompletely, connective splitting or not, sides glabrous, sutures papillate; staminode: 8–9 mm, included, 0.3–0.5 mm diam., tip straight, distal 4–5 mm sparsely papillate, papillae violet, to 0.2 mm; style 10–12 mm. |
7–12 × 4–6 mm, glabrous. |
3.5–6.5 mm. |
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Flowering May–Jun. |
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Sand dunes, ponderosa pine forests, mixed shrublands. |
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1800–2200 m. (5900–7200 ft.) |
AZ; CA; NM; UT; n Mexico
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UT |
Varieties 2 (2 in the flora). Two varieties, distinguished on the basis of inflorescence indument, are largely allopatric. Penstemon ×crideri A. Nelson, a putative hybrid between P. eatonii and P. pseudospectabilis, has been reported from Arizona (A. Nelson 1936). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon ammophilus is known from Garfield, Kane, and Washington counties. Populations occur on aeolian sand, derived from the Navajo Sandstone. The fistulose stems of P. ammophilus are unique in Penstemon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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FNA vol. 17, p. 253. |
FNA vol. 17, p. 161. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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M. E. Jones: Contr. W. Bot. 12: 66. (1908) — (as Pentstemon) |
N. H. Holmgren & L. M. Shultz: Brittonia 34: 381, fig. 1. (1982) |
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