Chelan beardtongue, Chelan penstemon, Chilean beardtongue
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Wilcox's beardtongue, Wilcox's penstemon
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Herbs. |
Herbs. |
ascending or erect, 8–40(–60) cm, glabrate, retrorsely hairy, or glandular-pubescent, not glaucous. |
ascending to erect, (20–)35–100 cm, glabrous or sparsely to densely puberulent, sometimes also sparsely glandular-pubescent, especially distally, not glaucous. |
basal and cauline, not leathery, retrorsely hairy or pubescent, sometimes also glandular-pubescent; basal and proximal cauline 25–100(–150) × (4–)9–20(–34) mm, blade elliptic to ovate, base tapered, margins ± serrate to dentate, apex obtuse to acute; cauline 2–5 pairs, sessile, 11–60 × 5–25(–30) mm, blade triangular-ovate to lanceolate, base truncate to cordate-clasping, margins serrate to dentate, apex acute, rarely obtuse. |
basal and cauline, ± leathery or not, glabrous or puberulent to pubescent, especially along midveins; basal and proximal cauline (25–)35–190 × (5–)12–55 mm, blade triangular to ovate or lanceolate, base tapered, rarely truncate, margins ± serrate or dentate, apex acute, rarely obtuse; cauline 2–5 pairs, short-petiolate or sessile, 35–90 × (8–)15–50 mm, blade ovate to oblong or lanceolate, base tapered to clasping or cordate-clasping, margins serrate to dentate, apex obtuse to acute. |
interrupted, cylindric, 4–28 cm, axis densely glandular-pubescent, rarely sparsely glandular-pubescent, verticillasters 3–8, cymes 3–11-flowered, 2 per node; proximal bracts ovate to lanceolate, 8–58 × 4–25 mm, margins serrate, sometimes entire; peduncles and pedicels ascending to erect, densely glandular-pubescent. |
interrupted, cylindric to conic, (5–)8–30 cm, axis sparsely to moderately glandular-pubescent, verticillasters 3–7, cymes 3–11-flowered, 2 per node; proximal bracts ovate to lanceolate, (8–)18–75 × (5–)8–35 mm, margins ± serrate to dentate; peduncles and pedicels spreading to ascending, ± glandular-pubescent. |
calyx lobes ovate to lanceolate, 2.5–4.5(–6) × 0.9–1.5 mm, glandular-pubescent; corolla blue to lavender, purple, or violet, with purple nectar guides, funnelform, 11–16 mm, glandular-pubescent externally, sparsely white-villous internally abaxially, tube 4–5 mm, throat gradually inflated, 3–6 mm diam., rounded abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, navicular to subexplanate, 0.6–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 7–9 mm, included or slightly exserted, 0.2–0.3 mm diam., tip recurved, distal 0.5–1 mm densely pilose, hairs yellow, to 0.8 mm; style 8–10 mm. |
calyx lobes ovate, 2.4–5.5 × 0.9–1.8 mm, margins broadly scarious, glandular-pubescent; corolla blue to violet or light blue, with reddish purple nectar guides, funnelform, 13–23 mm, glandular-pubescent externally, sparsely white- or yellow-villous internally abaxially, tube 4–8 mm, throat gradually inflated, 4–9 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular to subexplanate, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or ± papillate; staminode 8–11 mm, reaching orifice, 0.2–0.3 mm diam., tip straight to recurved, distal 1–5 mm pilose to villous, hairs yellow, to 1.2 mm; style 9–13 mm. |
4–7 × 3–4 mm, glabrous. |
4–6.5 × 3–4 mm, glabrous. |
= 16. |
= 16, 32. |
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Flowering Apr–Jul. |
Flowering May–Aug. |
Open grassy, sandy, gravelly, and rocky slopes, pine and pine-fir woodlands, sagebrush shrublands. |
Rocky or gravelly slopes in forests and woodlands, roadcuts. |
300–2000 m. (1000–6600 ft.) |
300–2300 m. (1000–7500 ft.) |
WA; BC
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ID; MT; OR; WA
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Penstemon pruinosus occurs along the eastern slope of the Cascade Range in Chelan, Douglas, Grant, and Kittitas counties, Washington, east into the scablands of Adams and Franklin counties, Washington, and north into extreme southern British Columbia. A blue dye is made from the flowers of Penstemon pruinosus by the Okanagan-Coville tribe of northeastern Washington (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon wilcoxii hybridizes with P. albertinus in areas of sympatry in east-central Idaho and west-central Montana (D. D. Keck 1945; A. Cronquist 1959; D. V. Clark 1971). Putative hybrids and/or introgressants exhibit varying degrees of morphologic intermediacy between the parent species. Many herbarium specimens are difficult to determine with confidence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 218. |
FNA vol. 17, p. 227. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wrightii, P. xylus, P. yampaënsis |
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P. ovatus var. pinetorum |
Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1280. (1829) — (as Pentstemon pruinosum) |
Rydberg: Bull. Torrey Bot. Club 28: 28. (1901) — (as Pentstemon) |
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