Blue Mountain beardtongue, Blue Mountain penstemon, Blue Mountains beardtongue, Pennell's beardtongue, Pennell's penstemon
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Deam's beardtongue
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Herbs. |
ascending to erect, 18–85 cm, glabrous, not glaucous. |
ascending to erect, 40–90 cm, retrorsely hairy proximally, usually retrorsely hairy distally, not glaucous. |
basal and cauline, ± leathery or not, glabrous, not glaucous; basal and proximal cauline 55–250 × 7–40 mm, blade lanceolate to elliptic, base tapered, margins entire, apex obtuse to acute; cauline 3–5 pairs, short-petiolate or sessile, 40–83 × 10–36 mm, blade ovate to lanceolate, base tapered to clasping, apex obtuse to acute. |
basal and cauline, basal sometimes withering by anthesis, not leathery, glabrate to puberulent; basal and proximal cauline 40–150 × 10–26 mm, blade oval to elliptic, base tapered, margins entire or ± denticulate, apex rounded to obtuse; cauline 7 or 8 pairs, sessile, 17–110 × 3–20 mm, blade oblanceolate to lanceolate, base tapered to clasping, margins entire or denticulate to serrulate, apex obtuse to acute. |
continuous, cylindric, 4–35 cm, axis glabrous, verticillasters 4–11, cymes 1–6-flowered, 2 per node; proximal bracts ovate to lanceolate, 22–85 × 11–35 mm; peduncles and pedicels glabrous, peduncles to 20 mm, pedicels 1–6 mm. |
interrupted, conic, 11–22 cm, axis glabrous or sparsely glandular-pubescent, verticillasters 4–6, cymes (2–)5–8-flowered, 2 per node; proximal bracts lanceolate, 10–15 × 3–5 mm, margins entire; peduncles and pedicels ascending to erect, glabrous or sparsely glandular-pubescent. |
calyx lobes ovate to lanceolate, (5.3–)6.5–8.5 × 1.6–3 mm, apex acuminate to caudate, glabrous; corolla blue to violet or purple, with reddish purple nectar guides, ventricose, 29–35 mm, glabrous externally, glabrous internally, tube 8–11 mm, throat gradually inflated, not constricted at orifice, 9–10 mm diam., slightly 2-ridged abaxially; stamens: longer pair reaching orifice, pollen sacs divergent to opposite, sigmoid, 2–2.7 mm, dehiscing incompletely, proximal 1/5–1/3 indehiscent, connective not splitting, sides sparsely hirsute, hairs white or tan, to 0.1 mm, especially near connective, sometimes merely scurfy near connective, sutures denticulate, teeth to 0.1 mm; staminode 16–20 mm, reaching orifice, 0.5–1 mm diam., tip straight, distal 4–6 mm ± uniformly sparsely pilose, hairs yellow, to 1 mm; style 20–24 mm. |
calyx lobes ovate, 1.5–3.5(–4) × 0.9–1.5 mm, sparsely glandular-pubescent; corolla white, sometimes tinged lavender or purple, with reddish purple nectar guides, ventricose, 15–20(–22) mm, glandular-pubescent externally, sparsely white-pubescent internally abaxially, tube 5–6 mm, throat abruptly inflated, 5–7 mm diam., slightly 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sometimes sparsely pubescent, hairs white, to 0.2 mm, sutures papillate; staminode 8–10 mm, reaching orifice, 0.3–0.4 mm diam., tip straight to slightly recurved, distal 3–4 mm pilose, hairs yellowish, to 1 mm; style 10–12 mm. |
10–13 × 8–10 mm. |
5–8 × 4–5 mm, glabrous. |
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Flowering May–Jun. |
Flowering May–Jun. |
Rocky ridges, sandy to rocky slopes, coniferous forests. |
Rocky oak woods, fields. |
1300–1800 m. (4300–5900 ft.) |
100–200 m. (300–700 ft.) |
OR; WA
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IN |
Penstemon pennellianus occurs primarily in the Blue Mountains of northeastern Oregon (Grant, Umatilla, Union, and Wallowa counties) and southeastern Washington (Asotin, Columbia, and Garfield counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A. C. Koelling (1964) assessed Penstemon deamii and its allies using morphological, cytological, and breeding studies. He concluded that P. deamii is a distinct member of the eastern North American polyploids that include P. calycosus, P. digitalis, and P. laevigatus. Geographically sympatric with P. calycosus and P. digitalis, P. deamii is distinguished from those two species by its shorter calyx lobes [1.5–3.5(–4) mm versus 4–9 mm], corollas [15–20(–22) mm versus (17–)20–35 mm], and pollen sacs (0.8–1 mm versus 1.2–1.7 mm). The species appears to be limited to Clark, Crawford, Floyd, Harrison, and Marion counties (K. Yatskievych 2000). Reports of P. deamii from southern Illinois appear to be based on specimens of questionable provenance or misidentifications (Koelling). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 176. |
FNA vol. 17, p. 199. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
D. D. Keck: Amer. Midl. Naturalist 23: 614. (1940) |
Pennell: Monogr. Acad. Nat. Sci. Philadelphia 1: 212. (1935) |
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