eastern white beardtongue, pale beardtongue, pale penstemon
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Gibbens' beardtongue
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Herbs. |
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ascending to erect, 25–55(–65) cm, retrorsely hairy and glandular-villous proximally, retrorsely hairy and glandular-pubescent distally, glandular hairs sometimes sparse, not glaucous. |
ascending to erect, 10–37 cm, pubescent to retrorsely hairy, sometimes glabrous proximally, glandular-pubescent distally, not glaucous. |
basal and cauline, basal often withering by anthesis, not leathery, sparsely to densely pubescent, sometimes also with scattered glandular hairs, abaxially, sparsely pubescent adaxially; basal and proximal cauline 20–120(–180) × 5–35(–40) mm, blade spatulate to obovate or ovate, base tapered, margins subentire or ± serrate or dentate, apex rounded to obtuse or acute; cauline 4–8 pairs, sessile, 22–100 × 4–24 mm, blade ovate to lanceolate, base truncate to clasping, margins subentire or ± serrate or dentate, apex acute to acuminate. |
basal and cauline, or basal absent or reduced, not leathery, proximals glabrous or puberulent to scabrous, distals puberulent or scabrous to glandular-pubescent, not glaucous; basal and proximal cauline 15–90 × 2–7(–8) mm, blade oblanceolate to linear, base tapered, margins entire, apex obtuse to acute; cauline 6–10 pairs, short-petiolate or sessile, 9–68 × 1–5 mm, blade oblanceolate to linear, base tapered to truncate, apex obtuse to acute. |
interrupted, conic, 5–26(–30) cm, axis sparsely to densely glandular-pubescent, verticillasters 3–8, cymes 2–8(–16)-flowered, 2 per node; proximal bracts lanceolate, 7–72 × 2–22 mm, margins entire or ± dentate; peduncles and pedicels ascending, sparsely to densely glandular-pubescent. |
interrupted, secund, (2–)5–14 cm, axis glandular-pubescent, verticillasters (2–)5–8, cymes 1–3-flowered, 1 or 2 per node; proximal bracts lanceolate, proximals 8–43 × 1–3 mm; peduncles and pedicels glandular-pubescent. |
calyx lobes ovate to lanceolate, (2–)3–5 × 1–3 mm, sparsely glandular-pubescent; corolla white, sometimes tinged lavender or blue, with reddish purple nectar guides, tubular to tubular-funnelform, 16–22 mm, glandular-pubescent externally, moderately whitish or yellowish lanate internally abaxially, tube 4–6 mm, throat slightly inflated, 4–7 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, (0.8–)1–1.2 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 10–12 mm, exserted, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 8–9 mm moderately to densely villous, hairs yellow or golden yellow, to 1.5 mm; style 12–14 mm. |
calyx lobes ovate to lanceolate, 3.5–7(–8) × 1.8–2.5 mm, glandular-pubescent; corolla lavender to light blue or light violet, with faint reddish purple nectar guides, funnelform, (15–)16–18(–20) mm, glandular-pubescent externally, sparsely to moderately glandular-pubescent or glandular internally abaxially, tube 5.5–6.5 mm, throat gradually inflated, not constricted at orifice, 5–6 mm diam., rounded abaxially; stamens: longer pair reaching orifice, pollen sacs divergent, navicular, 1–1.4 mm, dehiscing incompletely, proximal 1/5 indehiscent, connective not splitting, sides moderately hirsute, hairs white, to 0.7 mm, sutures denticulate, teeth to 0.1 mm; staminode 8–10 mm, included, 0.6–0.7 mm diam., tip recurved, distal 1–3 mm sparsely pilose or lanate, hairs yellow or whitish, to 1 mm; style 11–16 mm. |
5–7 × 3–5 mm, glabrous. |
5–8 × 3.5–5 mm. |
= 16. |
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Flowering Apr–Jun(–Jul). |
Flowering Jun–Sep. |
Tallgrass prairie, limestone and sandstone glades, barrens, rocky oak-hickory woodlands. |
Barren hills, pinyon-juniper woodlands, sagebrush and greasewood-saltbush shrublands. |
10–300 m. (0–1000 ft.) |
1700–2300 m. (5600–7500 ft.) |
AL; AR; CT; DC; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; ON
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CO; UT; WY |
Penstemon pallidus is concentrated in the Central Lowlands and Ozark Plateau of the central United States. F. W. Pennell (1935) believed it to be native there, having spread eastward as a result of human activities, as in Ontario. Penstemon brevisepalus, which has been synonymized with P. pallidus by many authors, is treated here as a distinct species. Characters distinguishing the species are discussed under 147. P. brevisepalus. Some specimens from north-central Arkansas and north-central Missouri are morphologically intermediate between Penstemon pallidus and P. arkansanus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon gibbensii is known from fewer than ten populations in Moffat and Rio Blanco counties, Colorado, Daggett County, Utah, and Carbon and Sweetwater counties, Wyoming (S. L. O’Kane 1988; E. C. Neese and N. D. Atwood 2003). Plants occur on shales or sandstones of the Browns Park Formation and Green River Formation (B. L. Heidel 2009). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 214. |
FNA vol. 17, p. 167. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
Small: Fl. S.E. U.S., 1060, 1337. (1903) — (as Pentstemon) |
Dorn: Brittonia 34: 334. (1982) |
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