eastern white beardtongue, pale beardtongue, pale penstemon
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scarlet bugler
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Herbs. |
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ascending to erect, 25–55(–65) cm, retrorsely hairy and glandular-villous proximally, retrorsely hairy and glandular-pubescent distally, glandular hairs sometimes sparse, not glaucous. |
ascending to erect, 30–120 cm, glaucous. |
basal and cauline, basal often withering by anthesis, not leathery, sparsely to densely pubescent, sometimes also with scattered glandular hairs, abaxially, sparsely pubescent adaxially; basal and proximal cauline 20–120(–180) × 5–35(–40) mm, blade spatulate to obovate or ovate, base tapered, margins subentire or ± serrate or dentate, apex rounded to obtuse or acute; cauline 4–8 pairs, sessile, 22–100 × 4–24 mm, blade ovate to lanceolate, base truncate to clasping, margins subentire or ± serrate or dentate, apex acute to acuminate. |
glabrous, glaucous; cauline 5–11 pairs, short-petiolate or sessile, 40–100 × 10–40 mm, blade ovate to lanceolate, base tapered to auriculate-clasping, margins entire, apex rounded to acute. |
interrupted, conic, 5–26(–30) cm, axis sparsely to densely glandular-pubescent, verticillasters 3–8, cymes 2–8(–16)-flowered, 2 per node; proximal bracts lanceolate, 7–72 × 2–22 mm, margins entire or ± dentate; peduncles and pedicels ascending, sparsely to densely glandular-pubescent. |
interrupted, secund to ± cylindric, 15–60(–100) cm, axis glabrous or obscurely scabrous, verticillasters 8–18(–22), cymes 1–5(–11)-flowered; proximal bracts ovate to lanceolate, (3–)7–25(–87) × (1–)3–14(–30) mm; peduncles and pedicels erect, glabrous or obscurely scabrous. |
calyx lobes ovate to lanceolate, (2–)3–5 × 1–3 mm, sparsely glandular-pubescent; corolla white, sometimes tinged lavender or blue, with reddish purple nectar guides, tubular to tubular-funnelform, 16–22 mm, glandular-pubescent externally, moderately whitish or yellowish lanate internally abaxially, tube 4–6 mm, throat slightly inflated, 4–7 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, (0.8–)1–1.2 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 10–12 mm, exserted, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 8–9 mm moderately to densely villous, hairs yellow or golden yellow, to 1.5 mm; style 12–14 mm. |
calyx lobes ovate to lanceolate, 3–6 × 2–3 mm, margins erose, glabrous; corolla scarlet, without nectar guides, nearly radially symmetric, weakly bilabiate, salverform, 25–33 mm, glabrous externally, glabrous internally, tube 8–10 mm, throat slightly inflated, 4.5–6 mm diam., rounded abaxially; stamens: longer pair reaching orifice, pollen sacs explanate, 0.9–1.2 mm, sutures smooth; staminode 13–14 mm, terete to slightly flattened distally, 0.1–0.3 mm diam., tip straight, glabrous; style 18–27 mm, exserted. |
5–7 × 3–5 mm, glabrous. |
8–11 × 6–7 mm. |
= 16. |
= 16. |
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Flowering Apr–Jun(–Jul). |
Flowering Feb–Jul. |
Tallgrass prairie, limestone and sandstone glades, barrens, rocky oak-hickory woodlands. |
Hillsides, chaparral, oak, pinyon-juniper, and Joshua tree woodlands, coastal sage scrub, pine forests. |
10–300 m. (0–1000 ft.) |
0–1800 m. (0–5900 ft.) |
AL; AR; CT; DC; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; ON
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CA; Mexico (Baja California)
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Penstemon pallidus is concentrated in the Central Lowlands and Ozark Plateau of the central United States. F. W. Pennell (1935) believed it to be native there, having spread eastward as a result of human activities, as in Ontario. Penstemon brevisepalus, which has been synonymized with P. pallidus by many authors, is treated here as a distinct species. Characters distinguishing the species are discussed under 147. P. brevisepalus. Some specimens from north-central Arkansas and north-central Missouri are morphologically intermediate between Penstemon pallidus and P. arkansanus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon ×parishii A. Gray, a naturally occurring hybrid between P. centranthifolius and P. spectabilis (A. D. Wolfe and W. J. Elisens 1993), has been reported from Los Angeles, Riverside, San Bernardino, and San Diego counties, California (D. D. Keck 1937; Paul Wilson and M. Valenzuela 2002). A wild hybrid between P. centranthifolius and P. eatonii also was reported from San Bernardino County, California (Wilson and Valenzuela). Penstemon ×dubius Davidson was described from Mount Lowe in the San Gabriel Mountains, Los Angeles County, California, growing with P. centranthifolius and P. grinnellii. Hybridization between P. centranthifolius and P. grinnellii has been documented using allozyme and DNA data (Wolfe and Elisens 1993, 1994). Three southern and central Californian tribes of Native Americans used Penstemon centranthifolius for drugs, food, and decorations (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 214. |
FNA vol. 17, p. 151. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Chelone centranthifolia |
Small: Fl. S.E. U.S., 1060, 1337. (1903) — (as Pentstemon) |
(Bentham) Bentham: Scroph. Ind., 7. (1835) — (as Pentstemon centranthifolium) |
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