thick-leaf beardtongue, thickleaf penstemon
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Susanville beardtongue, volcanic beardtongue
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Subshrubs. |
ascending to erect, (10–)15–57 cm, glabrous. |
ascending to erect, 18–70 cm, glandular-pubescent, sometimes densely so distally, not glaucous. |
basal and cauline, glabrous; basal and proximal cauline 18–180 × 5–45(–53) mm, blade spatulate to lanceolate, base tapered, apex rounded to obtuse, acute, or mucronate; cauline 2–5 pairs, short-petiolate or sessile, 15–80 × 7–32 mm, blade obovate, ovate, oblanceolate, or lanceolate, base tapered, clasping, or cordate-clasping, apex acute to rounded or mucronate. |
cauline, opposite or subopposite, not leathery, glandular-pubescent; cauline (4–)7–10 pairs, sessile or proximals short-petiolate, 14–60 × 8–20(–30) mm, blade obovate to ovate or lanceolate, base tapered to clasping, margins dentate, dentate-serrate, or serrate, apex acute to acuminate. |
continuous or interrupted, cylindric, (5–)9–27 cm, axis glabrous, verticillasters (2–)4–10, cymes (1–)3–6-flowered; proximal bracts ovate to lanceolate or oblong, 9–46 × 7–15 mm; peduncles and pedicels glabrous, peduncles to 26 mm. |
interrupted or continuous, cylindric, 13–35 cm, axis densely glandular-pubescent, verticillasters 9–18, cymes 1–5-flowered, 2 per node; proximal bracts ovate to lanceolate, (7–)15–40 × (2–)8–18 mm, margins dentate to dentate-serrate; peduncles and pedicels ascending to erect, densely glandular-pubescent. |
calyx lobes ovate to lanceolate, (2.5–)4–7 × 1.8–2.8 mm, margins entire or erose, broadly scarious, apex acute to acuminate or caudate, glabrous; corolla blue to violet, sometimes lavender, with or without reddish violet nectar guides, tubular-funnelform to ventricose, 12–20 mm, glabrous externally, glabrous internally or sparsely white-villous abaxially, tube 4–6 mm, throat gradually inflated, 5–6 mm diam., rounded abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, (0.7–)1–1.5 mm, sutures smooth or papillate; staminode 8–12 mm, reaching orifice or slightly exserted, (0.2–)0.5–2 mm diam., tip recurved, prominently bifurcate, distal 4–8 mm densely villous, hairs yellow to golden yellow, to 2.5 mm; style 11–15 mm. |
calyx lobes ovate to lanceolate, 3–6 × 0.8–1.6 mm, densely glandular-pubescent; corolla white to light lavender or light pink, with reddish violet nectar guides, funnelform, 9–11 mm, glandular-pubescent externally, glandular-pubescent internally, tube 3–4 mm, throat slightly inflated, 3–4 mm diam., rounded abaxially; stamens included, pollen sacs opposite, navicular to subexplanate, 0.5–0.7 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 9–10 mm, included or exserted, 0.2–0.3 mm diam., tip straight, glabrous; style 7–8 mm. |
10–14 × 5–7 mm. |
4–5 × 2.5–3 mm, glabrous. |
2–3 mm. |
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Flowering Jun–Jul. |
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Igneous soils and crevices, basalt in sagebrush shrublands and pine woodlands. |
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1200–1800 m. (3900–5900 ft.) |
AZ; NV; UT
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CA; NV |
Varieties 2 (2 in the flora). E. C. Neese (1993) considered Penstemon mucronatus transitional to P. pachyphyllus var. pachyphyllus in northeastern Utah. Penstemon mucronatus is distinguished from P. pachyphyllus var. pachyphyllus by its narrower staminodes bearing shorter, less tangled hairs, and its nectar guides extending onto the corolla limbs. The Havasupai tribe of northwestern Arizona uses leaves of Penstemon pachyphyllus to make a deer call for hunting (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon sudans is known from northeastern California (Lassen, Modoc, and Placer counties); it has been reported for Washoe County, Nevada. D. D. Keck (1940b) observed that some plants of P. deustus from southeastern Oregon (Curry, Jackson, and Josephine counties) approach P. sudans in glandularity, though he considered these to be one extreme of the morphologic spectrum exhibited by P. deustus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Staminodes 8–9 × 1–2 mm, distal 6–8 mm densely villous, hairs to 2.5 mm; Utah. | var. pachyphyllus |
1. Staminodes 10–12 × (0.2–)0.5–1 mm, distal 4–7 mm densely villous, hairs to 1.5 mm; Arizona, Nevada, Utah. | var. congestus |
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FNA vol. 17, p. 123. |
FNA vol. 17, p. 223. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. nitidus var. major |
P. deustus subsp. sudans |
A. Gray ex Rydberg: Fl. Rocky Mts., 770, 1066. (1917) — (as Pentstemon) |
M. E. Jones: Contr. W. Bot. 8: 37. (1898) — (as Pentstemon) |
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