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thick-leaf beardtongue, thickleaf penstemon

Apache beardtongue

Habit Herbs.
Stems

ascending to erect, (10–)15–57 cm, glabrous.

erect, (6–)10–45 cm, retrorsely hairy, also glandular-pubescent distally, not glaucous.

Leaves

basal and cauline, glabrous;

basal and proximal cauline 18–180 × 5–45(–53) mm, blade spatulate to lanceolate, base tapered, apex rounded to obtuse, acute, or mucronate;

cauline 2–5 pairs, short-petiolate or sessile, 15–80 × 7–32 mm, blade obovate, ovate, oblanceolate, or lanceolate, base tapered, clasping, or cordate-clasping, apex acute to rounded or mucronate.

basal and cauline, not leathery, proximals usually retrorsely hairy proximally, especially along midveins and margins, mostly glabrate distally, distals retrorsely hairy and, sometimes, sparsely glandular-pubescent, sometimes ± glaucescent;

basal and proximal cauline 15–75 × (2–)5–19 mm, blade spatulate to oblanceolate or elliptic, base tapered, margins entire, apex rounded to obtuse or acute, sometimes mucronate;

cauline 2–5 pairs, sessile, 11–70 × (1–)4–15 mm, blade of proximal leaves oblanceolate to lanceolate, distal leaves lanceolate, base tapered to clasping, margins entire, rarely ± denticulate distally, apex acute to acuminate.

Thyrses

continuous or interrupted, cylindric, (5–)9–27 cm, axis glabrous, verticillasters (2–)4–10, cymes (1–)3–6-flowered;

proximal bracts ovate to lanceolate or oblong, 9–46 × 7–15 mm;

peduncles and pedicels glabrous, peduncles to 26 mm.

interrupted, secund, (1–)3–24(–30) cm, axis glandular-pubescent, verticillasters (2 or)3–7, cymes 1–4-flowered, (1 or)2 per node;

proximal bracts lanceolate to linear, 7–38(–60) × 1–7 mm, margins entire;

peduncles and pedicels erect, glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, (2.5–)4–7 × 1.8–2.8 mm, margins entire or erose, broadly scarious, apex acute to acuminate or caudate, glabrous;

corolla blue to violet, sometimes lavender, with or without reddish violet nectar guides, tubular-funnelform to ventricose, 12–20 mm, glabrous externally, glabrous internally or sparsely white-villous abaxially, tube 4–6 mm, throat gradually inflated, 5–6 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, (0.7–)1–1.5 mm, sutures smooth or papillate;

staminode 8–12 mm, reaching orifice or slightly exserted, (0.2–)0.5–2 mm diam., tip recurved, prominently bifurcate, distal 4–8 mm densely villous, hairs yellow to golden yellow, to 2.5 mm;

style 11–15 mm.

calyx lobes lanceolate, 2.8–6 × 0.9–2 mm, glandular-pubescent;

corolla blue to violet-blue, with reddish purple nectar guides, tubular to tubular-funnelform, 11–20 mm, glandular-pubescent externally, sparsely to moderately white-villous to white-lanate internally abaxially, tube 5–7 mm, throat slightly inflated, 3–6 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, navicular, 0.8–1.2 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 10–12 mm, included or slightly exserted, 0.3–0.4 mm diam., tip straight to slightly recurved, distal 3–6 mm densely pilose, hairs golden, to 2 mm;

style 9–13 mm.

Capsules

10–14 × 5–7 mm.

5–7 × 3–4.5 mm, glabrous.

Seeds

2–3 mm.

2n

= 16.

Penstemon pachyphyllus

Penstemon oliganthus

Phenology Flowering Jun–Aug(–Oct).
Habitat Montane meadows, ciénagas, clearings in pine and spruce-fir forests.
Elevation 2400–3500 m. (7900–11500 ft.)
Distribution
from FNA
AZ; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

E. C. Neese (1993) considered Penstemon mucronatus transitional to P. pachyphyllus var. pachyphyllus in northeastern Utah. Penstemon mucronatus is distinguished from P. pachyphyllus var. pachyphyllus by its narrower staminodes bearing shorter, less tangled hairs, and its nectar guides extending onto the corolla limbs.

The Havasupai tribe of northwestern Arizona uses leaves of Penstemon pachyphyllus to make a deer call for hunting (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon oliganthus is known from the Mogollon Rim (Coconino County) and White Mountains of east-central Arizona (Apache and Greenlee counties) and the Jemez and San Mateo mountains of northwestern New Mexico (McKinley, Sandoval, and Valencia counties). Crosswhite described P. pseudoparvus from five specimens from the Magdalena and San Mateo mountains in Socorro County, New Mexico, and more than 24 specimens of P. oliganthus from Arizona and New Mexico. He separated the two species based on stem indument (obscurely puberulent in P. oliganthus versus obviously puberulent in P. pseudoparvus), and flower orientation and shape (horizontal or more usually drooping and little inflated in P. oliganthus versus ascending and not inflated in P. pseudoparvus). More than 60 collections referable to these species were examined for this treatment, including the types, specimens annotated by Crosswhite, and 20 specimens from the San Mateo and Magdalena mountains in Socorro County; the two taxa appear to be indistinguishable.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Staminodes 8–9 × 1–2 mm, distal 6–8 mm densely villous, hairs to 2.5 mm; Utah.
var. pachyphyllus
1. Staminodes 10–12 × (0.2–)0.5–1 mm, distal 4–7 mm densely villous, hairs to 1.5 mm; Arizona, Nevada, Utah.
var. congestus
Source FNA vol. 17, p. 123. FNA vol. 17, p. 213.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. pachyphyllus var. congestus, P. pachyphyllus var. pachyphyllus
Synonyms P. nitidus var. major P. pseudoparvus
Name authority A. Gray ex Rydberg: Fl. Rocky Mts., 770, 1066. (1917) — (as Pentstemon) Wooton & Standley: Contr. U.S. Natl. Herb. 16: 172. (1913) — (as Pentstemon)
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