thick-leaf beardtongue, thickleaf penstemon
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Oklahoma beardtongue
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Herbs. |
ascending to erect, (10–)15–57 cm, glabrous. |
ascending to erect, (15–)35–55 cm, spreading-puberulent or retrorsely hairy proximally, glandular-pubescent medially and distally, not glaucous. |
basal and cauline, glabrous; basal and proximal cauline 18–180 × 5–45(–53) mm, blade spatulate to lanceolate, base tapered, apex rounded to obtuse, acute, or mucronate; cauline 2–5 pairs, short-petiolate or sessile, 15–80 × 7–32 mm, blade obovate, ovate, oblanceolate, or lanceolate, base tapered, clasping, or cordate-clasping, apex acute to rounded or mucronate. |
basal and cauline, basal often withering by anthesis, not leathery, puberulent or retrorsely hairy; basal and proximal cauline 25–80(–110) × 5–24(–32) mm, blade spatulate to obovate, oblanceolate, or lanceolate, base tapered, margins subentire or ± denticulate or serrate, apex rounded to obtuse or acute; cauline 2–9 pairs, short-petiolate or sessile, (25–)60–120 × 4–20 mm, blade lanceolate to linear, base tapered to clasping, margins entire or ± denticulate to serrate, apex acute. |
continuous or interrupted, cylindric, (5–)9–27 cm, axis glabrous, verticillasters (2–)4–10, cymes (1–)3–6-flowered; proximal bracts ovate to lanceolate or oblong, 9–46 × 7–15 mm; peduncles and pedicels glabrous, peduncles to 26 mm. |
interrupted, cylindric, (5–)8–18(–25) cm, axis glandular-pubescent, verticillasters (2 or)3–6, cymes 2–4(–6)-flowered, 2 per node; proximal bracts lanceolate to linear, 60 × 6 mm, margins entire; peduncles and pedicels ascending to erect, glandular-pubescent. |
calyx lobes ovate to lanceolate, (2.5–)4–7 × 1.8–2.8 mm, margins entire or erose, broadly scarious, apex acute to acuminate or caudate, glabrous; corolla blue to violet, sometimes lavender, with or without reddish violet nectar guides, tubular-funnelform to ventricose, 12–20 mm, glabrous externally, glabrous internally or sparsely white-villous abaxially, tube 4–6 mm, throat gradually inflated, 5–6 mm diam., rounded abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, (0.7–)1–1.5 mm, sutures smooth or papillate; staminode 8–12 mm, reaching orifice or slightly exserted, (0.2–)0.5–2 mm diam., tip recurved, prominently bifurcate, distal 4–8 mm densely villous, hairs yellow to golden yellow, to 2.5 mm; style 11–15 mm. |
calyx lobes ovate to lanceolate, 5–7 × 3–4 mm, glandular-pubescent; corolla white to ochroleucous, without nectar guides, personate, tubular, 24–32 mm, glandular-pubescent externally, densely yellowish lanate internally abaxially, tube slightly differentiated from throat, 6–10 mm, throat slightly inflated, 6–8 mm diam., 2-ridged abaxially; stamens included, pollen sacs divergent, navicular to subexplanate, 1–1.5 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 18–20 mm, included or slightly exserted, 0.4–0.6 mm diam., tip straight, distal 14–16 mm ± villous, hairs yellowish orange, to 1.5 mm; style 20–22 mm. |
10–14 × 5–7 mm. |
8–13 × 3–5 mm, glabrous. |
2–3 mm. |
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= 16. |
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Flowering Apr–Jun. |
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Tallgrass prairies, open woods. |
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200–300 m. (700–1000 ft.) |
AZ; NV; UT
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OK; TX |
Varieties 2 (2 in the flora). E. C. Neese (1993) considered Penstemon mucronatus transitional to P. pachyphyllus var. pachyphyllus in northeastern Utah. Penstemon mucronatus is distinguished from P. pachyphyllus var. pachyphyllus by its narrower staminodes bearing shorter, less tangled hairs, and its nectar guides extending onto the corolla limbs. The Havasupai tribe of northwestern Arizona uses leaves of Penstemon pachyphyllus to make a deer call for hunting (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon oklahomensis is known from 26 counties primarily in east-central Oklahoma and in Lamar County, Texas (J. N. Mink et al. 2010). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Staminodes 8–9 × 1–2 mm, distal 6–8 mm densely villous, hairs to 2.5 mm; Utah. | var. pachyphyllus |
1. Staminodes 10–12 × (0.2–)0.5–1 mm, distal 4–7 mm densely villous, hairs to 1.5 mm; Arizona, Nevada, Utah. | var. congestus |
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FNA vol. 17, p. 123. |
FNA vol. 17, p. 212. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. nitidus var. major |
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A. Gray ex Rydberg: Fl. Rocky Mts., 770, 1066. (1917) — (as Pentstemon) |
Pennell: Monogr. Acad. Nat. Sci. Philadelphia 1: 237. (1935) |
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