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derived penstemon, Plumas County beardtongue

Cusick's beardtongue, Cusick's penstemon

Habit Subshrubs. Herbs or subshrubs.
Stems

ascending to erect, 20–80 cm, glabrous, sometimes ± puberulent, hairs pointed, glaucous.

ascending to erect, 15–45 cm, retrorsely hairy, hairs pointed, not glaucous.

Leaves

basal and cauline, basal sometimes few, opposite, glabrous, sometimes proximals puberulent, hairs pointed, glaucous;

basal and proximal cauline 15–80 × 2–10 mm, blade spatulate to oblanceolate, base tapered, margins entire, apex rounded to obtuse or acute;

cauline 4–6 pairs, sessile, 20–80 × 3–9 mm, blade ovate to lanceolate, base tapered, margins entire, apex acute.

cauline, opposite or subalternate, retrorsely hairy, hairs pointed, not glaucous;

cauline 4–8 pairs, short-petiolate or sessile, 16–70 × 1–8(–14) mm, blade lanceolate to linear, base tapered, margins entire, apex obtuse to acute.

Thyrses

interrupted, cylindric, 5–25 cm, axis glandular-pubescent, verticillasters 4–6, cymes 1–3-flowered, 2 per node;

proximal bracts ovate to lanceolate, 4–50 × 1–8 mm;

peduncles and pedicels ascending, glandular-pubescent.

interrupted, cylindric, 4–15 cm, axis puberulent or retrorsely hairy, verticillasters 3–10, cymes 1–5(–7)-flowered, 2 per node;

proximal bracts lanceolate to linear, 7–50 × 1–6 mm;

peduncles and pedicels ascending to erect, puberulent or retrorsely hairy.

Flowers

calyx lobes ovate to lanceolate, 3.8–6 × 1.5–2.2 mm, glandular-pubescent;

corolla violet to blue or purple, without nectar guides, ventricose to ventricose-ampliate, 25–38 mm, glandular-pubescent externally, glabrous internally, tube 7–9 mm, throat gradually inflated, 8–10 mm diam., 2-ridged abaxially;

stamens included, filaments glabrous, pollen sacs parallel, 2.4–3.2 mm, distal 1/5–1/3 indehiscent, sides pilose to lanate, hairs white, to 0.6 mm near filament attachment, sutures denticulate, teeth to 0.2 mm;

staminode 11–15 mm, included, 0.4–0.6 mm diam., glabrous;

style 15–20 mm.

calyx lobes ovate, 3–5.5 × 1.4–2.3 mm, glabrous or sparsely puberulent;

corolla blue to violet or purple, without nectar guides, ampliate, 16–21 mm, glabrous externally, glabrous internally, tube 4–5 mm, throat gradually inflated, 5–6 mm diam., slightly 2-ridged abaxially;

stamens included, filaments glabrous, pollen sacs parallel, 1–1.5(–1.8) mm, distal 1/2–2/3 indehiscent, sides hispidulous, hairs white, to 0.1 mm near filament attachment, sutures denticulate, teeth to 0.1 mm;

staminode 10–12 mm, reaching orifice or barely exserted, 0.5–0.7 mm diam., glabrous;

style 11–13 mm.

Capsules

7–9 × 4–5 mm.

6–7 × 4.5–6 mm.

2n

= 64.

= 16.

Penstemon neotericus

Penstemon cusickii

Phenology Flowering May–Aug(–Oct). Flowering May–Jul.
Habitat Volcanic soils, scrub, open pine and fir forests. Rocky, often basaltic, sagebrush shrublands.
Elevation 1000–2200 m. (3300–7200 ft.) 600–1500 m. (2000–4900 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon neotericus is known from the Cascade Range and northern Sierra Nevada of northern California (Butte, Lassen, Placer, Plumas, Shasta, Sierra, Siskiyou, Tehama, and Yuba counties). D. D. Keck (1932) used morphologic and geographic data to hypothesize that P. neotericus, an octoploid, arose through hybridization between P. azureus and P. laetus. J. Clausen (1933) provided cytological evidence supporting that hypothesis. A diploid chromosome number of 2n = 32 listed by N. H. Holmgren (1993) is probably a transcription error.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon cusickii is known from west-central and southwestern Idaho (Ada, Blaine, Canyon, Elmore, Gem, Owyhee, and Washington counties) and eastern Oregon (Baker, Grant, Harney, Malheur, Union, and Wheeler counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 238. FNA vol. 17, p. 232.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority D. D. Keck: Univ. Calif. Publ. Bot. 16: 398, fig. 13. (1932) A. Gray: Proc. Amer. Acad. Arts 16: 106. (1880) — (as Pentstemon)
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