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derived penstemon, Plumas County beardtongue

long-sepal beardtongue

Habit Subshrubs. Herbs.
Stems

ascending to erect, 20–80 cm, glabrous, sometimes ± puberulent, hairs pointed, glaucous.

erect, (40–)60–120 cm, puberulent, at least proximally, slightly glaucous or not.

Leaves

basal and cauline, basal sometimes few, opposite, glabrous, sometimes proximals puberulent, hairs pointed, glaucous;

basal and proximal cauline 15–80 × 2–10 mm, blade spatulate to oblanceolate, base tapered, margins entire, apex rounded to obtuse or acute;

cauline 4–6 pairs, sessile, 20–80 × 3–9 mm, blade ovate to lanceolate, base tapered, margins entire, apex acute.

basal and cauline, basal often withering by anthesis, not leathery, glabrous;

basal and proximal cauline 47–110 × 10–24 mm, blade oblanceolate, base tapered, margins entire or ± serrate, apex rounded to obtuse;

cauline 3–7 pairs, sessile or proximals short-petiolate, (14–)44–155 × (2–)13–46 mm, blade lanceolate to ovate, proximals sometimes oblanceolate, base clasping or tapered, margins serrate, apex acute to acuminate.

Thyrses

interrupted, cylindric, 5–25 cm, axis glandular-pubescent, verticillasters 4–6, cymes 1–3-flowered, 2 per node;

proximal bracts ovate to lanceolate, 4–50 × 1–8 mm;

peduncles and pedicels ascending, glandular-pubescent.

interrupted, conic, (5–)14–20 cm, axis sparsely puberulent and glandular-pubescent, rarely glabrous, verticillasters 2–5, cymes (1–)5–15-flowered, 2 per node;

proximal bracts ovate to lanceolate, sometimes linear, 13–80 × (2–)5–32 mm, margins serrate, sometimes entire;

peduncles and pedicels spreading to ascending, sparsely glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 3.8–6 × 1.5–2.2 mm, glandular-pubescent;

corolla violet to blue or purple, without nectar guides, ventricose to ventricose-ampliate, 25–38 mm, glandular-pubescent externally, glabrous internally, tube 7–9 mm, throat gradually inflated, 8–10 mm diam., 2-ridged abaxially;

stamens included, filaments glabrous, pollen sacs parallel, 2.4–3.2 mm, distal 1/5–1/3 indehiscent, sides pilose to lanate, hairs white, to 0.6 mm near filament attachment, sutures denticulate, teeth to 0.2 mm;

staminode 11–15 mm, included, 0.4–0.6 mm diam., glabrous;

style 15–20 mm.

calyx lobes lanceolate, 5–9 × 0.8–1.9 mm, apex acuminate, sparsely glandular-pubescent;

corolla pale lavender to violet, with faint violet nectar guides, ventricose, 20–35 mm, glandular-pubescent externally, ± white-pubescent internally abaxially, tube 5–7 mm, throat abruptly inflated, 8–11 mm diam., slightly 2-ridged abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, navicular, 1.2–1.4 mm, dehiscing completely, connective splitting, sides glabrous, rarely sparsely pubescent, hairs white, to 0.2 mm, sutures papillate;

staminode 11–13 mm, reaching orifice, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 2–5 mm sparsely pubescent, hairs yellowish, to 1 mm;

style 14–16 mm.

Capsules

7–9 × 4–5 mm.

7–8 × 4–5 mm, glabrous.

2n

= 64.

= 96.

Penstemon neotericus

Penstemon calycosus

Phenology Flowering May–Aug(–Oct). Flowering May–Jun.
Habitat Volcanic soils, scrub, open pine and fir forests. Woodlands, meadows, rocky slopes, stream banks.
Elevation 1000–2200 m. (3300–7200 ft.) 10–300 m. (0–1000 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DC; GA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NJ; NY; OH; PA; RI; SC; TN; VA; VT; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon neotericus is known from the Cascade Range and northern Sierra Nevada of northern California (Butte, Lassen, Placer, Plumas, Shasta, Sierra, Siskiyou, Tehama, and Yuba counties). D. D. Keck (1932) used morphologic and geographic data to hypothesize that P. neotericus, an octoploid, arose through hybridization between P. azureus and P. laetus. J. Clausen (1933) provided cytological evidence supporting that hypothesis. A diploid chromosome number of 2n = 32 listed by N. H. Holmgren (1993) is probably a transcription error.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon calycosus is concentrated in the Central Lowlands and Interior Low Plateaus; occurrences farther south and east, especially in New England, may be from introductions (F. W. Pennell 1935). The range lies almost entirely within that of P. digitalis, but historic ranges of both P. calycosus and P. digitalis are difficult to delimit due to both past and continuing introduction and spread of both species. Calyx lobe lengths to 12 mm are reported in the literature; those appear to be from fruiting calyces. Penstemon calycosus usually has glabrous anthers, but many specimens, especially from Illinois and Ohio, bear hairs on the adaxial surfaces of the anthers. Some populations from this region exhibit variation in vegetative (leaf size and bract shape and margins) and floral characters (calyx lobe shape and size, corolla color, and anther pubescence), suggesting possible hybridization with P. digitalis, an observation that would be consistent with the findings of A. C. Koelling (1964).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 238. FNA vol. 17, p. 197.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority D. D. Keck: Univ. Calif. Publ. Bot. 16: 398, fig. 13. (1932) Small: Bull. Torrey Bot. Club 25: 470. (1898) — (as Pentstemon)
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