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derived penstemon, Plumas County beardtongue

Nevada sand-dune beardtongue

Habit Subshrubs. Herbs.
Caudex

rhizomelike.

Stems

ascending to erect, 20–80 cm, glabrous, sometimes ± puberulent, hairs pointed, glaucous.

ascending to erect, 10–27 cm, glabrous, glutinous, usually covered with sand, not glaucous.

Leaves

basal and cauline, basal sometimes few, opposite, glabrous, sometimes proximals puberulent, hairs pointed, glaucous;

basal and proximal cauline 15–80 × 2–10 mm, blade spatulate to oblanceolate, base tapered, margins entire, apex rounded to obtuse or acute;

cauline 4–6 pairs, sessile, 20–80 × 3–9 mm, blade ovate to lanceolate, base tapered, margins entire, apex acute.

cauline, leathery, glabrous, sometimes glutinous;

cauline 3–5 pairs, short-petiolate or sessile, (12–)20–50(–55) × (3–)10–24 mm, blade obovate or spatulate proximally, obovate to lanceolate distally, base tapered, margins dentate, apex rounded to obtuse or acute.

Thyrses

interrupted, cylindric, 5–25 cm, axis glandular-pubescent, verticillasters 4–6, cymes 1–3-flowered, 2 per node;

proximal bracts ovate to lanceolate, 4–50 × 1–8 mm;

peduncles and pedicels ascending, glandular-pubescent.

continuous, cylindric, (1–)2–9 cm, axis glabrous or sparsely puberulent, glutinous, verticillasters 2–8, cymes 2–6-flowered, 2 per node;

proximal bracts lanceolate, 16–58 × 3–12 mm, margins dentate;

peduncles and pedicels erect, glabrous or puberulent, glutinous.

Flowers

calyx lobes ovate to lanceolate, 3.8–6 × 1.5–2.2 mm, glandular-pubescent;

corolla violet to blue or purple, without nectar guides, ventricose to ventricose-ampliate, 25–38 mm, glandular-pubescent externally, glabrous internally, tube 7–9 mm, throat gradually inflated, 8–10 mm diam., 2-ridged abaxially;

stamens included, filaments glabrous, pollen sacs parallel, 2.4–3.2 mm, distal 1/5–1/3 indehiscent, sides pilose to lanate, hairs white, to 0.6 mm near filament attachment, sutures denticulate, teeth to 0.2 mm;

staminode 11–15 mm, included, 0.4–0.6 mm diam., glabrous;

style 15–20 mm.

calyx lobes ovate to lanceolate, 4.5–8.4 × 1.6–3 mm, glutinous;

corolla white to lavender, with reddish purple nectar guides, tubular-funnelform, 11–15(–18) mm, glabrous externally, sparsely white-pilose internally abaxially, tube 6–7 mm, throat gradually inflated, 3–5 mm diam., rounded abaxially;

stamens included, pollen sacs opposite, navicular to subexplanate, 0.8–0.9 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth;

staminode 7–10 mm, included, 0.6–0.8 mm diam., tip slightly recurved, distal 0.5–2.5 mm sparsely pilose, hairs white, to 0.5 mm;

style 7–8 mm.

Capsules

7–9 × 4–5 mm.

5–8 × 4–5 mm, glabrous.

2n

= 64.

= 16.

Penstemon neotericus

Penstemon arenarius

Phenology Flowering May–Aug(–Oct). Flowering May–Jul.
Habitat Volcanic soils, scrub, open pine and fir forests. Loose sands, dunes, valleys, desert shrublands.
Elevation 1000–2200 m. (3300–7200 ft.) 1200–1800 m. (3900–5900 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
NV
[BONAP county map]
Discussion

Penstemon neotericus is known from the Cascade Range and northern Sierra Nevada of northern California (Butte, Lassen, Placer, Plumas, Shasta, Sierra, Siskiyou, Tehama, and Yuba counties). D. D. Keck (1932) used morphologic and geographic data to hypothesize that P. neotericus, an octoploid, arose through hybridization between P. azureus and P. laetus. J. Clausen (1933) provided cytological evidence supporting that hypothesis. A diploid chromosome number of 2n = 32 listed by N. H. Holmgren (1993) is probably a transcription error.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon arenarius has been documented in Churchill, Mineral, and Nye counties. Plants usually are covered with sand because of their glutinous herbage.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 238. FNA vol. 17, p. 191.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority D. D. Keck: Univ. Calif. Publ. Bot. 16: 398, fig. 13. (1932) Greene: Pittonia 1: 282. (1889) — (as Pentstemon)
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