cup-leaf beardtongue, scarlet beardtongue
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eastern white beardtongue, pale beardtongue, pale penstemon
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Herbs. |
ascending to erect, 50–100(–150) cm, glabrous. |
ascending to erect, 25–55(–65) cm, retrorsely hairy and glandular-villous proximally, retrorsely hairy and glandular-pubescent distally, glandular hairs sometimes sparse, not glaucous. |
basal and cauline, glabrous; basal and proximal cauline 40–100 × (8–)14–30 mm, blade spatulate to ovate, base tapered, apex rounded to obtuse, rarely retuse; cauline 5–9 pairs, sessile, 12–110 × 16–70 mm, blade oblong to ovate, base clasping to connate-perfoliate, apex obtuse to acute. |
basal and cauline, basal often withering by anthesis, not leathery, sparsely to densely pubescent, sometimes also with scattered glandular hairs, abaxially, sparsely pubescent adaxially; basal and proximal cauline 20–120(–180) × 5–35(–40) mm, blade spatulate to obovate or ovate, base tapered, margins subentire or ± serrate or dentate, apex rounded to obtuse or acute; cauline 4–8 pairs, sessile, 22–100 × 4–24 mm, blade ovate to lanceolate, base truncate to clasping, margins subentire or ± serrate or dentate, apex acute to acuminate. |
interrupted, cylindric, 15–50 cm, axis glabrous, verticillasters 6–10(–16), cymes 1–5-flowered; proximal bracts ovate, 9–45 × 14–40 mm; peduncles and pedicels glabrous. |
interrupted, conic, 5–26(–30) cm, axis sparsely to densely glandular-pubescent, verticillasters 3–8, cymes 2–8(–16)-flowered, 2 per node; proximal bracts lanceolate, 7–72 × 2–22 mm, margins entire or ± dentate; peduncles and pedicels ascending, sparsely to densely glandular-pubescent. |
calyx lobes lanceolate, 4.5–7 × 1.8–2.9 mm, margins entire, narrowly scarious, glabrous; corolla red, without nectar guides, tubular-funnelform, 32–36 mm, glabrous externally, glabrous internally, tube 8–11 mm, throat abruptly inflated, 9–12 mm diam., rounded abaxially; stamens exserted, pollen sacs opposite, explanate, 1.3–1.7 mm, sutures smooth; staminode 18–20 mm, exserted, 0.2–0.3 mm diam., tip straight to recurved, glabrous; style 24–31 mm. |
calyx lobes ovate to lanceolate, (2–)3–5 × 1–3 mm, sparsely glandular-pubescent; corolla white, sometimes tinged lavender or blue, with reddish purple nectar guides, tubular to tubular-funnelform, 16–22 mm, glandular-pubescent externally, moderately whitish or yellowish lanate internally abaxially, tube 4–6 mm, throat slightly inflated, 4–7 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, (0.8–)1–1.2 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 10–12 mm, exserted, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 8–9 mm moderately to densely villous, hairs yellow or golden yellow, to 1.5 mm; style 12–14 mm. |
9–13 × 6–10 mm. |
5–7 × 3–5 mm, glabrous. |
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= 16. |
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Flowering Mar–Jun. |
Flowering Apr–Jun(–Jul). |
Sandy soils, deciduous and pine woodlands, sandhill prairies. |
Tallgrass prairie, limestone and sandstone glades, barrens, rocky oak-hickory woodlands. |
10–200 m. (0–700 ft.) |
10–300 m. (0–1000 ft.) |
AR; LA; OK; TX
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AL; AR; CT; DC; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; ON
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F. S. Crosswhite and C. D. Crosswhite (1981) hypothesized that Penstemon murrayanus was derived from P. grandiflorus, with which it shares many morphologic features. The two species have been crossed artificially, yielding hybrids that once were commercially popular (G. Viehmeyer 1958; R. Nold 1999). Molecular data support the sister relationship of P. murrayanus and P. grandiflorus (C. A. Wessinger et al. 2016). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon pallidus is concentrated in the Central Lowlands and Ozark Plateau of the central United States. F. W. Pennell (1935) believed it to be native there, having spread eastward as a result of human activities, as in Ontario. Penstemon brevisepalus, which has been synonymized with P. pallidus by many authors, is treated here as a distinct species. Characters distinguishing the species are discussed under 147. P. brevisepalus. Some specimens from north-central Arkansas and north-central Missouri are morphologically intermediate between Penstemon pallidus and P. arkansanus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 121. |
FNA vol. 17, p. 214. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
Hooker: Bot. Mag. 63: plate 3472. (1836) — (as Pentstemon) |
Small: Fl. S.E. U.S., 1060, 1337. (1903) — (as Pentstemon) |
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