cup-leaf beardtongue, scarlet beardtongue
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Amargosa beardtongue, Death Valley beardtongue, Death Valley penstemon
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Shrubs or subshrubs. |
ascending to erect, 50–100(–150) cm, glabrous. |
ascending to erect, 30–60 cm, glaucous. |
basal and cauline, glabrous; basal and proximal cauline 40–100 × (8–)14–30 mm, blade spatulate to ovate, base tapered, apex rounded to obtuse, rarely retuse; cauline 5–9 pairs, sessile, 12–110 × 16–70 mm, blade oblong to ovate, base clasping to connate-perfoliate, apex obtuse to acute. |
cauline 6–12 pairs, sessile or short-petiolate, (12–)25–65 × 2–8 mm, blade lanceolate to linear, sometimes oblanceolate, base truncate or tapered, margins entire, rarely ± serrulate distally, apex obtuse to acute. |
interrupted, cylindric, 15–50 cm, axis glabrous, verticillasters 6–10(–16), cymes 1–5-flowered; proximal bracts ovate, 9–45 × 14–40 mm; peduncles and pedicels glabrous. |
interrupted, cylindric, 8–30 cm, axis glabrous, verticillasters 3–8, cymes 1–3-flowered; proximal bracts lanceolate to linear, 9–28 × 2–5 mm; peduncles and pedicels spreading to ascending, glabrous. |
calyx lobes lanceolate, 4.5–7 × 1.8–2.9 mm, margins entire, narrowly scarious, glabrous; corolla red, without nectar guides, tubular-funnelform, 32–36 mm, glabrous externally, glabrous internally, tube 8–11 mm, throat abruptly inflated, 9–12 mm diam., rounded abaxially; stamens exserted, pollen sacs opposite, explanate, 1.3–1.7 mm, sutures smooth; staminode 18–20 mm, exserted, 0.2–0.3 mm diam., tip straight to recurved, glabrous; style 24–31 mm. |
calyx lobes depressed-ovate to round or ovate, 3.5–6.5 × 2.5–5 mm, glabrous; corolla bluish lavender to lavender or light pink, with reddish purple nectar guides, strongly bilabiate, ventricose-ampliate, 22–28 mm, glabrous or glandular-pubescent externally, moderately to densely white-lanate internally abaxially, sometimes also sparsely to densely glandular-pubescent internally, tube 6–8 mm, length 1–1.5 times calyx lobes, throat abruptly inflated, constricted at orifice, 8–14 mm diam., 2-ridged abaxially; stamens included, filaments of shorter pair glandular-puberulent proximally, pollen sacs explanate to subexplanate, 1.6–2.2 mm, sutures smooth; staminode 15–19 mm, exserted, 0.6–0.8 mm diam., tip recurved, distal 8–9 mm moderately pilose to lanate, hairs yellow, to 2.5 mm; proximal 5–6 mm glandular-pubescent; style 19–24 mm, glabrous. |
9–13 × 6–10 mm. |
9–13 × 5–7 mm, glabrous. |
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= 16. |
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Flowering Mar–Jun. |
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Sandy soils, deciduous and pine woodlands, sandhill prairies. |
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10–200 m. (0–700 ft.) |
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AR; LA; OK; TX
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CA; NV
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F. S. Crosswhite and C. D. Crosswhite (1981) hypothesized that Penstemon murrayanus was derived from P. grandiflorus, with which it shares many morphologic features. The two species have been crossed artificially, yielding hybrids that once were commercially popular (G. Viehmeyer 1958; R. Nold 1999). Molecular data support the sister relationship of P. murrayanus and P. grandiflorus (C. A. Wessinger et al. 2016). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Penstemon fruticiformis is known from the Mojave Desert. Specimens from the Kingston Range and the eastern flank of the southern Sierra Nevada sometimes combine features of P. fruticiformis and P. incertus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Corollas glabrous externally; calyx lobes depressed-ovate to round or ovate, 3.5–5 × 3.2–5 mm. | var. fruticiformis |
1. Corollas ± glandular-pubescent externally; calyx lobes ovate, 4.5–6.5 × 2.5–4 mm. | var. amargosae |
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FNA vol. 17, p. 121. |
FNA vol. 17, p. 250. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Hooker: Bot. Mag. 63: plate 3472. (1836) — (as Pentstemon) |
Coville: Contr. U.S. Natl. Herb. 4: 170. (1893) — (as Pentstemon) |
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