mucronate penstemon, sheep creek beardtongue
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crested beardtongue, crested-tongue beardtongue, crested-tongue penstemon, fuzzy-tongue penstemon
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ascending to erect, 10–35(–46) cm, glabrous. |
ascending to erect, (2–)6–60 cm, retrorsely hairy, sometimes also villous or glandular-villous, or glabrate. |
basal and cauline, glabrous; basal and proximal cauline 23–65(–100) × (9–)13–40 mm, blade oblanceolate, base tapered, apex rounded to obtuse or acute; cauline (2–)4–6(–9) pairs, sessile, (8–)20–30 × 8–26 mm, blade ovate to orbiculate, base clasping, apex mucronate. |
basal and cauline, not leathery, glabrous or glabrate to retrorsely hairy and, sometimes, sparsely villous abaxially, glabrous or glabrate to retrorsely hairy and, sometimes, villous or glandular-villous adaxially; basal and proximal cauline petiolate, (14–)22–130(–180) × 3–30(–40) mm, blade spatulate to obovate, oblanceolate, elliptic, or linear, base tapered, margins entire or ± dentate to serrate, apex rounded to obtuse or acute; cauline 2–6 pairs, sessile or proximals short-petiolate, 18–90 × 3–20(–25) mm, blade oblanceolate to oblong, lanceolate, or linear, base tapered to clasping or cordate-clasping, margins entire or ± dentate to serrate, apex obtuse to acute. |
interrupted, cylindric, (2–)11–18(–27) cm, axis glabrous, verticillasters 6–9(–15), cymes 2–6-flowered; proximal bracts orbiculate to ovate, 7–28 × 3–26 mm; peduncles and pedicels glabrous. |
continuous or interrupted, cylindric, (1–)2–20(–27) cm, axis glandular-pubescent to glandular-villous, sometimes also retrorsely hairy, verticillasters 2–7(–9), cymes (1 or)2–6-flowered, 2 per node; proximal bracts oblanceolate to oblong or lanceolate, 12–75 × 2–23 mm; peduncles and pedicels glandular-villous or glandular-pubescent and, sometimes, also retrorsely hairy. |
calyx lobes ovate, (3–)4.5–6.5 × 1.5–2.3 mm, margins entire or erose, broadly scarious, glabrous; corolla blue to violet, with reddish violet nectar guides, guides sometimes passing onto limb, tubular-funnelform, (10–)13–17(–20) mm, glabrous externally, sparsely white-villous internally abaxially, tube (5–)7–10 mm, throat gradually inflated, 4.5–5.5(–7) mm diam., rounded abaxially; stamens included or longer pair reaching orifice, pollen sacs divergent to opposite, 0.7–1.1 mm, sutures papillate; staminode 7–13 mm, reaching orifice, (0.6–)0.9–1.9 mm diam., tip strongly recurved, distal 2–5 mm moderately to densely pilose, hairs brownish yellow or golden yellow, to 1 mm; style (7–)9–12 mm. |
calyx lobes lanceolate, 5.5–13 × 1.4–3 mm, glandular-pubescent to glandular-villous; corolla lavender to violet, purple, pink, or blue, with reddish purple nectar guides, funnelform or ventricose-ampliate, 16–35(–42) mm, sparsely to densely white- or yellow-villous internally abaxially, sometimes glandular-pubescent laterally, tube 4–9 mm, throat gradually to abruptly inflated, constricted or not at orifice, 6–14 mm diam., rounded abaxially; stamens included, pollen sacs widely divergent to opposite, navicular to subexplanate or explanate, 0.8–1.9 mm, dehiscing completely, sutures smooth or papillate; staminode 12–18 mm, exserted or prominently exserted, 0.3–0.9 mm diam., tip recurved, distal 6–15 mm sparsely to densely lanate, hairs yellow, yellowish, or orangish, to 4 mm, and medial hairs shorter, stiffer, and retrorse; style 9–20 mm, glabrous or proximal 1–10 mm sparsely glandular-pubescent. |
8–10 × 6–7 mm. |
6–13 × 4–6 mm, sparsely glandular-pubescent distally. |
= 16. |
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Flowering May–Jun. |
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Sandy or gravelly soils, juniper and pinyon-juniper woodlands. |
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1500–2500 m. (4900–8200 ft.) |
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CO; UT; WY
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CO; ID; MT; ND; NE; OR; SD; UT; WA; WY; AB; BC
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Penstemon mucronatus is known from Moffat and Rio Blanco counties, Colorado, Daggett and Uintah counties, Utah, and Carbon and Sweetwater counties, Wyoming. E. C. Neese (1993) considered it transitional to P. pachyphyllus var. pachyphyllus in northeastern Utah and treated it as a variety of that species. The two taxa often have similar leaf morphologies; P. mucronatus usually is easily distinguished from P. pachyphyllus var. pachyphyllus by its narrower staminode bearing shorter, less tangled hairs at the apex and nectar guides that extend onto the corolla limbs. Flowers of P. mucronatus are in many respects more similar to those of P. osterhoutii than they are to those of P. pachyphyllus. A chromosome count reported for P. pachyphyllus by F. S. Crosswhite and S. Kawano (1965) is referable to P. mucronatus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 5 (5 in the flora). D. D. Keck (1938) treated this complex as three species: Penstemon cleburnei, P. eriantherus (with three varieties), and P. whitedii (with three subspecies). Most authors followed A. Cronquist (1959), who employed a broader species concept; the approach by Cronquist is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Corollas constricted at orifices; styles 9–12(–14) mm; pollen sacs 0.8–1.2 mm. | var. cleburnei |
1. Corollas not constricted at orifices; styles 9–20 mm; pollen sacs 1–1.9 mm. | → 2 |
2. Stems glabrate or retrorsely hairy, rarely glandular-villous, proximally, glandular-villous distally; leaves glabrous or glabrate abaxially, glabrate or villous to glandular-villous adaxially. | var. whitedii |
2. Stems retrorsely hairy and, sometimes, sparsely villous, glandular-villous, or glandular-pubescent distally; leaves retrorsely hairy and, sometimes, sparsely villous or glabrous abaxially, retrorsely hairy or puberulent and, sometimes, villous or glandular-villous adaxially. | → 3 |
3. Corollas funnelform; staminodes sparsely lanate distally; leaves glabrous abaxially. | var. argillosus |
3. Corollas ventricose-ampliate; staminodes moderately to densely lanate distally; leaves retrorsely hairy, sparsely villous, or glabrous abaxially. | → 4 |
4. Pollen sacs explanate; stems retrorsely hairy and, usually, sparsely villous or glandular-villous distally; corollas (20–)22–35(–42) mm. | var. eriantherus |
4. Pollen sacs navicular to subexplanate; stems retrorsely hairy, sometimes also glandular-pubescent distally or wholly; corollas 18–25 mm. | var. redactus |
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FNA vol. 17, p. 121. |
FNA vol. 17, p. 133. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. pachyphyllus var. mucronatus |
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N. H. Holmgren: Brittonia 31: 234, fig. 10. (1979) |
Pursh: Fl. Amer. Sept. 2: 737. (1813) — (as Pentstemon erianthera) |
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