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toad-flax beardtongue, toadflax penstemon

Atwood's beardtongue, kaiparowits beardtongue

Stems

ascending to erect, 8–50 cm, retrorsely hairy, hairs pointed or white, scalelike.

ascending to erect, 14–35(–50) cm, glabrous or sparsely retrorsely hairy proximally, usually sparsely glandular-pubescent distally.

Leaves

not leathery, glabrous, glabrate, or sparsely to densely retrorsely hairy, hairs pointed, sometimes appressed, white, scalelike;

cauline 6–12(–20) pairs, petiolate or sessile, 4–26 × 0.5–3.5 mm, blade oblanceolate to lanceolate or linear, base tapered to truncate, apex mucronate.

basal and cauline, leathery or not, glabrous or sparsely retrorsely hairy proximally adaxially;

basal and proximal cauline petiolate, 30–80(–100) × 6–12 mm, blade oblanceolate to lanceolate, base tapered, margins entire or obscurely serrate, apex obtuse to acute or mucronate;

cauline 3 or 4 pairs, sessile, 28–78 × 3–7 mm, blade oblanceolate to lanceolate, base attenuate to clasping, margins entire, apex obtuse to acute.

Thyrses

continuous or interrupted, ± secund, 3–17 cm, axis glandular-pubescent, verticillasters 3–9(–12), cymes 1 or 2(or 3)-flowered, 1(or 2) per node;

proximal bracts lanceolate to linear, 6–25 × 0.7–1 mm;

peduncles and pedicels ascending to erect, retrorsely hairy, hairs pointed, and sparsely glandular-pubescent.

interrupted, cylindric, (3–)7–24 cm, axis glandular-pubescent, verticillasters 3–8, cymes 3–12-flowered, 2 per node;

proximal bracts lanceolate to linear, 22–65 × 2–10 mm;

peduncles and pedicels glandular-pubescent.

Flowers

calyx lobes ovate, rarely lanceolate, 4–7(–9) × 2–3 mm, broadly scarious-margined, apex acute to acuminate, glandular-pubescent, hairs pointed;

corolla blue or lavender, lined internally abaxially with reddish purple nectar guides, ventricose-ampliate, 16–20 mm, moderately white- or yellowish villous internally abaxially, tube 5–6 mm, throat abruptly inflated, 5.5–8 mm diam., slightly 2-ridged abaxially;

stamens: longer pair reaching orifice or slightly exserted, filaments glabrous, pollen sacs opposite, navicular, 0.9–1.3 mm, sutures papillate;

staminode 6–8 mm, exserted, flattened distally, 0.3–0.5 mm diam., tip straight to recurved, distal 1–5 mm densely pilose, hairs yellow or golden yellow, to 1.2 mm, rest of distal 1–7 mm glabrous or sparsely to moderately pilose;

style 10–11 mm.

calyx lobes lanceolate, 6–8(–9) × 1.5–2 mm, glandular-pubescent;

corolla blue to lavender, with dark violet nectar guides, bilabiate, tubular-funnelform, 10–13 mm, sparsely to moderately yellow-lanate internally abaxially, tube 4–5 mm, throat gradually inflated, not constricted at orifice, 3–5 mm diam., rounded abaxially;

stamens reaching orifice or longer pair exserted, pollen sacs opposite, navicular, 0.9–1.5 mm, dehiscing completely, sutures papillate;

staminode 6–8 mm, exserted, 0.3–0.4 mm diam., tip straight, distal 4–5 mm moderately to densely retrorsely-pubescent, hairs golden yellow, to 0.6 mm;

style 6–8 mm.

Capsules

5–9 × 3.5–5 mm.

8–12 × 4–7 mm.

Penstemon linarioides

Penstemon atwoodii

Phenology Flowering May–Jun.
Habitat Pinyon-juniper woodlands.
Elevation 1600–2400 m. (5200–7900 ft.)
Distribution
from FNA
AZ; CO; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Penstemon linarioides is widespread and highly variable. D. D. Keck (1937) recognized seven subspecies on the basis of habit, pubescence, leaf shape, and staminode bearding. Some of the variation on which those subspecies are based appears to be clinal or too variable within or among regional populations to be of taxonomic value.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

According to Welsh, Penstemon atwoodii is known only from the Kaiparowits Plateau of eastern Garfield and Kane counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves glabrous, glabrate, retrorsely hairy, or pubescent, hairs pointed.
var. sileri
1. Leaves sparsely to densely retrorsely hairy, hairs appressed, white, scalelike.
→ 2
2. Cauline leaf blades oblanceolate, distals sometimes lanceolate, 1.5–3.5 mm wide.
var. maguirei
2. Cauline leaf blades oblanceolate to linear, 0.5–2 mm wide.
→ 3
3. Staminodes: distal 3–5 mm densely pilose, hairs golden yellow or yellow, to 1 mm, rest of distal 1–4 mm sparsely to moderately pilose.
var. linarioides
3. Staminodes: distal 1 mm densely pilose, hairs yellow, to 0.8 mm, rest of distal 3–4 mm glabrous or sparsely pilose (with much shorter hairs).
var. coloradoensis
Source FNA vol. 17, p. 103. FNA vol. 17, p. 128.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. linarioides var. coloradoensis, P. linarioides var. linarioides, P. linarioides var. maguirei, P. linarioides var. sileri
Name authority A. Gray: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 112. (1859) — (as Pentstemon) S. L. Welsh: Great Basin Naturalist 35: 378. (1976)
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