toad-flax beardtongue, toadflax penstemon
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stiff-leaf beardtongue, stiffleaf penstemon
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Herbs. |
ascending to erect, 8–50 cm, retrorsely hairy, hairs pointed or white, scalelike. |
ascending to erect, 6–20(–25) cm, retrorsely hairy proximally, glabrous or retrorsely hairy distally, not glaucous. |
not leathery, glabrous, glabrate, or sparsely to densely retrorsely hairy, hairs pointed, sometimes appressed, white, scalelike; cauline 6–12(–20) pairs, petiolate or sessile, 4–26 × 0.5–3.5 mm, blade oblanceolate to lanceolate or linear, base tapered to truncate, apex mucronate. |
basal and cauline, not leathery, glabrous or ± puberulent; basal and proximal cauline 15–65 × 1–5(–7) mm, blade narrowly oblanceolate to linear, base tapered, margins entire, apex obtuse to acute; cauline 2–5 pairs, short-petiolate or sessile, 7–33 × 1–3 mm, blade lanceolate to linear, base sessile to clasping, margins entire, apex obtuse to acute. |
continuous or interrupted, ± secund, 3–17 cm, axis glandular-pubescent, verticillasters 3–9(–12), cymes 1 or 2(or 3)-flowered, 1(or 2) per node; proximal bracts lanceolate to linear, 6–25 × 0.7–1 mm; peduncles and pedicels ascending to erect, retrorsely hairy, hairs pointed, and sparsely glandular-pubescent. |
interrupted, cylindric, 2–11 cm, axis glandular-pubescent, sometimes also retrorsely hairy, verticillasters 2–6, cymes 1–5-flowered, 2 per node; proximal bracts lanceolate to linear, 9–32 × 1–6 mm, margins entire; peduncles and pedicels erect, ± glandular-pubescent. |
calyx lobes ovate, rarely lanceolate, 4–7(–9) × 2–3 mm, broadly scarious-margined, apex acute to acuminate, glandular-pubescent, hairs pointed; corolla blue or lavender, lined internally abaxially with reddish purple nectar guides, ventricose-ampliate, 16–20 mm, moderately white- or yellowish villous internally abaxially, tube 5–6 mm, throat abruptly inflated, 5.5–8 mm diam., slightly 2-ridged abaxially; stamens: longer pair reaching orifice or slightly exserted, filaments glabrous, pollen sacs opposite, navicular, 0.9–1.3 mm, sutures papillate; staminode 6–8 mm, exserted, flattened distally, 0.3–0.5 mm diam., tip straight to recurved, distal 1–5 mm densely pilose, hairs yellow or golden yellow, to 1.2 mm, rest of distal 1–7 mm glabrous or sparsely to moderately pilose; style 10–11 mm. |
calyx lobes ovate to lanceolate, 2.2–4.6 × 1–2.4 mm, glandular-pubescent; corolla blue to violet, with reddish purple guides, funnelform, 11–17 mm, glandular-pubescent externally, moderately whitish lanate internally abaxially, tube 3–4 mm, throat gradually inflated, 4–6 mm diam., 2-ridged abaxially; stamens included, pollen sacs divergent to opposite, subexplanate to explanate, 0.6–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 7–8 mm, reaching orifice, 0.2–0.4 mm diam., tip recurved, distal 0.5–1 mm densely pilose, hairs golden yellow, to 0.8 mm; style 9–10 mm. |
5–9 × 3.5–5 mm. |
5–7 × 4–5 mm, glabrous. |
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Flowering Jun–Jul. |
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Rocky hillsides, sagebrush shrublands, montane grasslands. |
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1500–2900 m. (4900–9500 ft.) |
AZ; CO; NM; NV; UT
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ID; MT; WY
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Varieties 4 (4 in the flora). Penstemon linarioides is widespread and highly variable. D. D. Keck (1937) recognized seven subspecies on the basis of habit, pubescence, leaf shape, and staminode bearding. Some of the variation on which those subspecies are based appears to be clinal or too variable within or among regional populations to be of taxonomic value. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon aridus is known from the Rocky Mountains in eastern Idaho and southwestern and south-central Montana and the Big Horn and northern Absaroka Mountains of Wyoming. The sometimes grasslike basal leaves distinguish P. aridus from P. humilis and P. virens. At lower elevations or in shaded, forest habitats, plants sometimes have broader basal leaves and tend to be much taller than typical plants; the broadly scarious-margined calyx lobes of P. aridus distinguish it from similar species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Leaves glabrous, glabrate, retrorsely hairy, or pubescent, hairs pointed. | var. sileri |
1. Leaves sparsely to densely retrorsely hairy, hairs appressed, white, scalelike. | → 2 |
2. Cauline leaf blades oblanceolate, distals sometimes lanceolate, 1.5–3.5 mm wide. | var. maguirei |
2. Cauline leaf blades oblanceolate to linear, 0.5–2 mm wide. | → 3 |
3. Staminodes: distal 3–5 mm densely pilose, hairs golden yellow or yellow, to 1 mm, rest of distal 1–4 mm sparsely to moderately pilose. | var. linarioides |
3. Staminodes: distal 1 mm densely pilose, hairs yellow, to 0.8 mm, rest of distal 3–4 mm glabrous or sparsely pilose (with much shorter hairs). | var. coloradoensis |
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FNA vol. 17, p. 103. |
FNA vol. 17, p. 192. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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A. Gray: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 112. (1859) — (as Pentstemon) |
Rydberg: Mem. New York Bot. Gard. 1: 348. (1900) — (as Pentstemon) |
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