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Leonard's beardtongue, Leonard's penstemon

blowout beardtongue

Habit Herbs or subshrubs. Plants: caudex rhizomelike.
Stems

decumbent to ascending or erect, (5–)10–30(–47) cm, puberulent or retrorsely hairy, hairs pointed, not glaucous.

decumbent to ascending, (15–)20–48 cm, glabrous.

Leaves

cauline, opposite, glabrous, sometimes proximals puberulent, hairs pointed, not glaucous;

cauline 2–7 pairs, short-petiolate or sessile, 11–48(–60) × 2–12 mm, blade oblanceolate to spatulate or obovate, rarely linear, base tapered, margins entire, apex rounded to obtuse or acute.

essentially cauline, basal absent or reduced, glabrous;

basal and proximal cauline (25–)55–130(–175) × 3–25 mm, blade linear to lanceolate, base tapered, apex acute to acuminate;

cauline 2–10 pairs, sessile, 60–110(–120) × 7–30 mm, blade lanceolate to linear, base clasping, apex acuminate to long-acuminate.

Thyrses

continuous or interrupted, cylindric to ± secund, (1–)2–15(–25) cm, axis glabrous or retrorsely hairy, verticillasters 2–8, cymes 1–4-flowered, 2 per node;

proximal bracts lanceolate, 10–35 × 1–4 mm;

peduncles and pedicels ascending to erect, peduncles glabrous or sparsely retrorsely hairy, pedicels glabrous or papillate distally.

continuous, cylindric, (2–)6–21(–34) cm, axis glabrous, verticillasters (2–)6–10(–17), cymes 1–8-flowered;

proximal bracts ovate, 58–120 × 20–45 mm;

peduncles and pedicels glabrous.

Flowers

calyx lobes ovate to lanceolate, 3–5.5 × 0.9–2.2 mm, margins entire or erose, scarious, apex acuminate to caudate, glabrous;

corolla lavender to blue or violet, with or without faint lavender nectar guides, funnelform to ventricose, 14–26 mm, glabrous externally, glabrous internally, tube 5–8 mm, throat gradually inflated, 6–8(–10) mm diam., slightly 2-ridged abaxially;

stamens included or longer pair reaching orifice, filaments glabrous, pollen sacs parallel, 0.9–1.9 mm, distal 1/2–2/3 indehiscent, sides glabrous or puberulent, hairs white, to 0.1 mm, sutures denticulate, teeth to 0.2 mm;

staminode 10–15 mm, included or reaching orifice, 0.6–0.9 mm diam., glabrous;

style 12–18 mm.

calyx lobes lanceolate to linear, 8–13 × 1–3 mm, margins entire or erose, herbaceous or scarious, glabrous;

corolla lavender to bluish, usually with magenta nectar guides, ampliate, 21–28 mm, glabrous externally, glabrous internally, tube 7–9 mm, throat abruptly inflated, 9–11 mm diam., rounded abaxially;

stamens included, pollen sacs opposite, 1.1–2 mm, sutures papillate;

staminode 13–16 mm, included, 1.2–3 mm diam., tip recurved, distal 0.1–5 mm sparsely to densely villous, hairs golden yellow, to 1 mm, rarely glabrous;

style 13–23 mm.

Capsules

6–8 × 2.5–4.5 mm.

8–12 × 5–9 mm.

2n

= 16.

Penstemon leonardii

Penstemon haydenii

Phenology Flowering May–Jul(–Sep).
Habitat Blowouts in sand dunes.
Elevation 900–2300 m. (3000–7500 ft.)
Distribution
from FNA
AZ; ID; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
NE; WY
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Penstemon leonardii is common in the Wasatch Mountains, Utah Plateaus, and mountain ranges in the eastern Basin and Range Province. Elements allied with it have been interpreted variously as one species comprising three varieties, two species (one with two varieties), or three species; the broadest concept is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon haydenii is known from the Nebraska Sandhills, where extant populations occur in Box Butte, Cherry, Garden, Hooker, Morrill, and Sheridan counties. Historic populations occurred in Thomas County (D. M. Sutherland 1988).

Penstemon haydenii was discovered in northern Carbon County, Wyoming, in the late 1990s some 300 km west of the nearest Nebraska populations; it might have been collected in Wyoming as early as 1877 (W. Fertig 2001). A morphometric analysis of Nebraska and Wyoming plants revealed differences that could justify recognition of Wyoming populations as a distinct variety (C. C. Freeman 2015). It is listed as endangered by the U.S. Department of the Interior.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas 20–26 mm; pollen sacs (1.2–)1.4–1.9 mm; e Nevada, w Utah.
var. patricus
1. Corollas 14–22 mm; pollen sacs 0.9–1.6 mm; nw Arizona, se Idaho, se Nevada, Utah.
→ 2
2. Corolla limbs blue to lavender; thyrses continuous; se Idaho, n Utah.
var. leonardii
2. Corolla limbs lavender to violet; thyrses continuous or interrupted; nw Arizona, se Nevada, sw Utah.
var. higginsii
Source FNA vol. 17, p. 237. FNA vol. 17, p. 119.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. leonardii var. higginsii, P. leonardii var. leonardii, P. leonardii var. patricus
Name authority Rydberg: Bull. Torrey Bot. Club 40: 483. (1913) — (as Pentstemon leonardi) S. Watson: Bot. Gaz. 16: 311. (1891) — (as Pentstemon haydeni)
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