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Leonard's beardtongue, Leonard's penstemon

Amargosa beardtongue, Death Valley beardtongue, Death Valley penstemon

Habit Herbs or subshrubs. Shrubs or subshrubs.
Stems

decumbent to ascending or erect, (5–)10–30(–47) cm, puberulent or retrorsely hairy, hairs pointed, not glaucous.

ascending to erect, 30–60 cm, glaucous.

Leaves

cauline, opposite, glabrous, sometimes proximals puberulent, hairs pointed, not glaucous;

cauline 2–7 pairs, short-petiolate or sessile, 11–48(–60) × 2–12 mm, blade oblanceolate to spatulate or obovate, rarely linear, base tapered, margins entire, apex rounded to obtuse or acute.

cauline 6–12 pairs, sessile or short-petiolate, (12–)25–65 × 2–8 mm, blade lanceolate to linear, sometimes oblanceolate, base truncate or tapered, margins entire, rarely ± serrulate distally, apex obtuse to acute.

Thyrses

continuous or interrupted, cylindric to ± secund, (1–)2–15(–25) cm, axis glabrous or retrorsely hairy, verticillasters 2–8, cymes 1–4-flowered, 2 per node;

proximal bracts lanceolate, 10–35 × 1–4 mm;

peduncles and pedicels ascending to erect, peduncles glabrous or sparsely retrorsely hairy, pedicels glabrous or papillate distally.

interrupted, cylindric, 8–30 cm, axis glabrous, verticillasters 3–8, cymes 1–3-flowered;

proximal bracts lanceolate to linear, 9–28 × 2–5 mm;

peduncles and pedicels spreading to ascending, glabrous.

Flowers

calyx lobes ovate to lanceolate, 3–5.5 × 0.9–2.2 mm, margins entire or erose, scarious, apex acuminate to caudate, glabrous;

corolla lavender to blue or violet, with or without faint lavender nectar guides, funnelform to ventricose, 14–26 mm, glabrous externally, glabrous internally, tube 5–8 mm, throat gradually inflated, 6–8(–10) mm diam., slightly 2-ridged abaxially;

stamens included or longer pair reaching orifice, filaments glabrous, pollen sacs parallel, 0.9–1.9 mm, distal 1/2–2/3 indehiscent, sides glabrous or puberulent, hairs white, to 0.1 mm, sutures denticulate, teeth to 0.2 mm;

staminode 10–15 mm, included or reaching orifice, 0.6–0.9 mm diam., glabrous;

style 12–18 mm.

calyx lobes depressed-ovate to round or ovate, 3.5–6.5 × 2.5–5 mm, glabrous;

corolla bluish lavender to lavender or light pink, with reddish purple nectar guides, strongly bilabiate, ventricose-ampliate, 22–28 mm, glabrous or glandular-pubescent externally, moderately to densely white-lanate internally abaxially, sometimes also sparsely to densely glandular-pubescent internally, tube 6–8 mm, length 1–1.5 times calyx lobes, throat abruptly inflated, constricted at orifice, 8–14 mm diam., 2-ridged abaxially;

stamens included, filaments of shorter pair glandular-puberulent proximally, pollen sacs explanate to subexplanate, 1.6–2.2 mm, sutures smooth;

staminode 15–19 mm, exserted, 0.6–0.8 mm diam., tip recurved, distal 8–9 mm moderately pilose to lanate, hairs yellow, to 2.5 mm;

proximal 5–6 mm glandular-pubescent;

style 19–24 mm, glabrous.

Capsules

6–8 × 2.5–4.5 mm.

9–13 × 5–7 mm, glabrous.

2n

= 16.

Penstemon leonardii

Penstemon fruticiformis

Distribution
from FNA
AZ; ID; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Penstemon leonardii is common in the Wasatch Mountains, Utah Plateaus, and mountain ranges in the eastern Basin and Range Province. Elements allied with it have been interpreted variously as one species comprising three varieties, two species (one with two varieties), or three species; the broadest concept is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Penstemon fruticiformis is known from the Mojave Desert. Specimens from the Kingston Range and the eastern flank of the southern Sierra Nevada sometimes combine features of P. fruticiformis and P. incertus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas 20–26 mm; pollen sacs (1.2–)1.4–1.9 mm; e Nevada, w Utah.
var. patricus
1. Corollas 14–22 mm; pollen sacs 0.9–1.6 mm; nw Arizona, se Idaho, se Nevada, Utah.
→ 2
2. Corolla limbs blue to lavender; thyrses continuous; se Idaho, n Utah.
var. leonardii
2. Corolla limbs lavender to violet; thyrses continuous or interrupted; nw Arizona, se Nevada, sw Utah.
var. higginsii
1. Corollas glabrous externally; calyx lobes depressed-ovate to round or ovate, 3.5–5 × 3.2–5 mm.
var. fruticiformis
1. Corollas ± glandular-pubescent externally; calyx lobes ovate, 4.5–6.5 × 2.5–4 mm.
var. amargosae
Source FNA vol. 17, p. 237. FNA vol. 17, p. 250.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. leonardii var. higginsii, P. leonardii var. leonardii, P. leonardii var. patricus
P. fruticiformis var. amargosae, P. fruticiformis var. fruticiformis
Name authority Rydberg: Bull. Torrey Bot. Club 40: 483. (1913) — (as Pentstemon leonardi) Coville: Contr. U.S. Natl. Herb. 4: 170. (1893) — (as Pentstemon)
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