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Leonard's beardtongue, Leonard's penstemon

pinto beardtongue, pinto penstemon, two-color beardtongue

Habit Herbs or subshrubs. Herbs.
Stems

decumbent to ascending or erect, (5–)10–30(–47) cm, puberulent or retrorsely hairy, hairs pointed, not glaucous.

ascending to erect, 60–150 cm, glaucous.

Leaves

cauline, opposite, glabrous, sometimes proximals puberulent, hairs pointed, not glaucous;

cauline 2–7 pairs, short-petiolate or sessile, 11–48(–60) × 2–12 mm, blade oblanceolate to spatulate or obovate, rarely linear, base tapered, margins entire, apex rounded to obtuse or acute.

basal and proximal cauline 37–110 × 10–50 mm, blade obovate, base tapered, margins coarsely serrate to coarsely dentate, apex obtuse to acute;

cauline 4–8 pairs, sessile, 40–110 × 15–55 mm, blade ovate, base connate-perfoliate, margins coarsely serrate to coarsely dentate, apex acute to acuminate.

Thyrses

continuous or interrupted, cylindric to ± secund, (1–)2–15(–25) cm, axis glabrous or retrorsely hairy, verticillasters 2–8, cymes 1–4-flowered, 2 per node;

proximal bracts lanceolate, 10–35 × 1–4 mm;

peduncles and pedicels ascending to erect, peduncles glabrous or sparsely retrorsely hairy, pedicels glabrous or papillate distally.

interrupted, secund, 16–90 cm, axis glandular-pubescent, verticillasters 9–23, cymes 1–4-flowered;

proximal bracts ovate to lanceolate, 8–25 × 3–28 mm;

peduncles and pedicels ascending to erect, glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 3–5.5 × 0.9–2.2 mm, margins entire or erose, scarious, apex acuminate to caudate, glabrous;

corolla lavender to blue or violet, with or without faint lavender nectar guides, funnelform to ventricose, 14–26 mm, glabrous externally, glabrous internally, tube 5–8 mm, throat gradually inflated, 6–8(–10) mm diam., slightly 2-ridged abaxially;

stamens included or longer pair reaching orifice, filaments glabrous, pollen sacs parallel, 0.9–1.9 mm, distal 1/2–2/3 indehiscent, sides glabrous or puberulent, hairs white, to 0.1 mm, sutures denticulate, teeth to 0.2 mm;

staminode 10–15 mm, included or reaching orifice, 0.6–0.9 mm diam., glabrous;

style 12–18 mm.

calyx lobes ovate, 4–6 × 2.2–3 mm, glandular-pubescent;

corolla pink to rose pink, purple, or light yellow, with or without reddish or reddish purple nectar guides, strongly bilabiate, ventricose, 18–27 mm, glandular-pubescent externally, glandular-pubescent internally, and sparsely white- or yellowish lanate internally abaxially, tube 7–10 mm, length 1.7–2 times calyx lobes, throat abruptly inflated, constricted at orifice, 6–9 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs explanate, 1.4–1.8 mm, sutures smooth;

staminode 14–16 mm, included or exserted, 0.3–0.4 mm diam., tip recurved, distal 4–5 mm lanate, hairs yellow, to 2.8 mm, proximal 3–4 mm glandular-puberulent;

style 14–16 mm, glabrous.

Capsules

6–8 × 2.5–4.5 mm.

10–13 × 5–7 mm, glabrous or sparsely glandular-pubescent distally.

Penstemon leonardii

Penstemon bicolor

Phenology Flowering Mar–Jun.
Habitat Gravelly soils, arroyos, roadsides, talus slopes, desert scrub, juniper woodlands.
Elevation 500–1700 m. (1600–5600 ft.)
Distribution
from FNA
AZ; ID; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Penstemon leonardii is common in the Wasatch Mountains, Utah Plateaus, and mountain ranges in the eastern Basin and Range Province. Elements allied with it have been interpreted variously as one species comprising three varieties, two species (one with two varieties), or three species; the broadest concept is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon bicolor is known from the Mojave Desert in the Black Mountains of northwestern Mohave County, Arizona, the Castle, Clark, and New York mountains of eastern San Bernardino County, California, and the desert mountain ranges of southern and western Clark County, Nevada. Two subspecies differing in corolla color have been recognized, with yellow-corolla forms restricted to Clark County, Nevada. Subspecies are not recognized here, following J. D. Morefield (2006).

Habitat destruction and genetic swamping caused by hybridization between Penstemon bicolor and P. palmeri threaten many populations of P. bicolor in the vicinity of Las Vegas, Nevada (G. Glenne 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas 20–26 mm; pollen sacs (1.2–)1.4–1.9 mm; e Nevada, w Utah.
var. patricus
1. Corollas 14–22 mm; pollen sacs 0.9–1.6 mm; nw Arizona, se Idaho, se Nevada, Utah.
→ 2
2. Corolla limbs blue to lavender; thyrses continuous; se Idaho, n Utah.
var. leonardii
2. Corolla limbs lavender to violet; thyrses continuous or interrupted; nw Arizona, se Nevada, sw Utah.
var. higginsii
Source FNA vol. 17, p. 237. FNA vol. 17, p. 247.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. leonardii var. higginsii, P. leonardii var. leonardii, P. leonardii var. patricus
Synonyms P. palmeri var. bicolor, P. bicolor subsp. roseus, P. pseudospectabilis subsp. bicolor
Name authority Rydberg: Bull. Torrey Bot. Club 40: 483. (1913) — (as Pentstemon leonardi) (Brandegee) Clokey & D. D. Keck: Bull. S. Calif. Acad. Sci. 38: 12. (1939)
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