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eastern beardtongue, eastern smooth beardtongue

Gorman's beardtongue, Gorman's penstemon

Habit Herbs.
Stems

ascending to erect, 40–115 cm, glabrous or retrorsely hairy, not glaucous.

ascending to erect, (6–)10–30(–65) cm, glabrous proximally, glandular-pubescent to glandular-villous distally.

Leaves

basal and cauline, basal sometimes withering by anthesis, not leathery, glabrous or puberulent;

basal and proximal cauline 40–150 × 7–48 mm, blade oblanceolate to lanceolate, base tapered, margins entire or ± serrate, apex obtuse to acute;

cauline 3–7 pairs, sessile or proximals sometimes petiolate, 18–128 × 8–28 mm, blade lanceolate, proximals sometimes oblanceolate, base clasping, sometimes tapered, margins entire or ± serrate, apex acute.

basal and cauline, not leathery, glabrous or distal cauline sometimes glabrous abaxially, glandular-pubescent adaxially;

basal and proximal cauline petiolate, (18–)35–100 × (2–)3–10 mm, blade spatulate to oblanceolate, base tapered, margins entire or obscurely serrate, apex rounded to obtuse or acute;

cauline 2–4 pairs, sessile, 15–68 × 2–16 mm, blade oblong to lanceolate, base tapered to clasping, margins entire or obscurely serrate, apex obtuse to acute.

Thyrses

interrupted, narrowly conic, (3–)8–20 cm, axis glabrous or retrorsely hairy and sparsely glandular-pubescent, verticillasters 2–6, cymes (2–)4–11-flowered, 2 per node;

proximal bracts lanceolate to linear, 8–38 × 1–3 mm, margins entire;

peduncles and pedicels spreading or ascending, glabrous or retrorsely hairy and sparsely glandular-pubescent.

continuous or ± interrupted, cylindric, (1–)3–15 cm, axis glandular-villous, verticillasters 2–6, cymes (1 or)2–5-flowered, 2 per node;

proximal bracts lanceolate to linear, (5–)10–56 × (1–)3–10 mm;

peduncles and pedicels glandular-pubescent.

Flowers

calyx lobes ovate, 3–6 × 1.5–2.1 mm, glabrous or glandular-pubescent;

corolla pale lavender to violet, with reddish purple nectar guides, ventricose, 15–22 mm, glandular-pubescent externally, sparsely to moderately white-lanate internally abaxially, tube 5–6 mm, throat abruptly inflated, 5–8 mm diam., slightly 2-ridged abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, navicular, 1–1.2 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 10–12 mm, exserted, 0.4–0.5 mm diam., tip straight to slightly recurved, distal 5–7 mm moderately to densely pubescent, hairs yellow, to 2.3 mm;

style 12–15 mm.

calyx lobes lanceolate, 7–10.5 × 1.4–2.2 mm, glandular-villous;

corolla lavender to purple or bluish violet, with reddish purple nectar guides, ventricose, 19–30 mm, sparsely white- or yellowish villous internally abaxially, tube 5–6 mm, throat abruptly inflated, not constricted at orifice, 8–14 mm diam., rounded abaxially;

stamens included, pollen sacs opposite, navicular, 1.3–1.5 mm, dehiscing completely, sutures smooth;

staminode 13–17 mm, prominently exserted, 0.4–0.6 mm diam., tip straight to recurved, distal 2–3 mm sparsely lanate, hairs yellow, to 2 mm, and medial 4–8 mm sparsely to moderately pilose, hairs yellow, to 1 mm;

style 11–14 mm.

Capsules

5–8 × 3.5–5 mm, glabrous.

7–10 × 4–5 mm.

2n

= 16.

Penstemon laevigatus

Penstemon gormanii

Phenology Flowering May–Jun. Flowering Jun–Aug.
Habitat Meadows, floodplain forests, fields, rock outcrops, calcareous bluffs. Rocky slopes, sand dunes, gravelly stream terraces, clearings in forests.
Elevation 10–400 m. (0–1300 ft.) 400–1200 m. (1300–3900 ft.)
Distribution
from FNA
AL; CT; DC; DE; FL; GA; KY; LA; MA; MD; MS; NC; NJ; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; BC; NT; YT
[BONAP county map]
Discussion

Penstemon laevigatus primarily is a species of the Piedmont and eastern Appalachians; scattered populations occur outside those regions. L. La Cour (1931) reported a chromosome count of n = 48 for P. laevigatus. Without a voucher, it is impossible to know if this count was for the species as treated here, or P. calycosus or P. digitalis, which have been treated as infraspecific taxa of P. laevigatus.

An infusion made from Penstemon laevigatus is used by the Cherokee tribe of the southeastern United States as a gastrointestinal aid (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon gormanii resembles P. eriantherus vars. argillosus and whitedii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 210. FNA vol. 17, p. 136.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority Aiton: Hort. Kew. 2: 361. (1789) — (as Pentstemon laevigata) Greene: Ottawa Naturalist 16: 39. (1902) — (as Pentstemon gormani)
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