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gay penstemon, mountain blue penstemon, western gray beardtongue

Canaan Mountain beardtongue, sandloving penstemon

Habit Herbs or subshrubs.
Stems

decumbent to ascending or erect, 15–90 cm, retrorsely hairy, hairs pointed, not glaucous.

decumbent to ascending, 5–26 cm, usually ± fistulose, glutinous and covered with sand, not glaucous.

Leaves

cauline, opposite, puberulent or retrorsely hairy, hairs pointed, sometimes glabrate, not glaucous;

cauline 4–8 pairs, sessile or proximals short-petiolate, 13–85(–110) × 2–22 mm, blade oblanceolate to lanceolate or linear, base tapered to truncate, margins entire, apex acute.

basal and cauline, not leathery, glutinous and covered with sand, not glaucous;

basal and proximal cauline 18–80 × 4–15 mm, blade oblanceolate, base tapered, margins entire, crisped, apex obtuse to acute;

cauline 2–5 pairs, sessile, 15–75 × 2–8 mm, blade oblanceolate to lanceolate, base tapered to truncate, margins crisped, apex rounded or obtuse to acute.

Thyrses

interrupted, cylindric, 10–30(–43) cm, axis glandular-pubescent, verticillasters 6–12, cymes 1–5-flowered, 2 per node;

proximal bracts lanceolate, rarely linear, 9–40 × 2–10 mm;

peduncles and pedicels spreading to ascending, glandular-pubescent.

continuous, cylindric, 4–15 cm, axis glutinous and covered with sand, verticillasters 4–10, cymes 1–6-flowered, 2 per node;

proximal bracts lanceolate, 14–45 × 2–5(–8) mm;

peduncles and pedicels glutinous and covered with sand.

Flowers

calyx lobes ovate to lanceolate, 4–11 × 1–2.8 mm, glandular-pubescent;

corolla violet to blue or purple, with or without violet nectar guides, funnelform to ampliate, 21–35 mm, glandular-pubescent externally, glabrous internally, tube 6–9 mm, throat gradually inflated, 8–10 mm diam., 2-ridged abaxially;

stamens included or longer pair reaching orifice, filaments glabrous, pollen sacs parallel, 1.8–2.8 mm, distal 1/5–1/3 indehiscent, sides pubescent to lanate, hairs white, to 1 mm near filament attachment, sutures denticulate, teeth to 0.3 mm;

staminode 17–19 mm, included or reaching orifice, 0.5–0.6 mm diam., glabrous;

style 16–22 mm.

calyx lobes lanceolate, 5–6.5 × 1–2 mm, glutinous and covered with sand;

corolla lavender to blue-lavender, with violet nectar guides, weakly ventricose, 14–17 mm, glabrous externally, usually glutinous and covered with sand distally, especially on lobes, glabrous internally, tube 5–6 mm, throat gradually inflated, not constricted at orifice, 5–6 mm diam., slightly 2-ridged abaxially;

stamens: longer pair reaching orifice, pollen sacs opposite, navicular, 0.9–1.4 mm, dehiscing completely or incompletely, connective splitting or not, sides glabrous, sutures papillate;

staminode: 8–9 mm, included, 0.3–0.5 mm diam., tip straight, distal 4–5 mm sparsely papillate, papillae violet, to 0.2 mm;

style 10–12 mm.

Capsules

7–9 × 4–6 mm.

3.5–6.5 mm.

Penstemon laetus

Penstemon ammophilus

Phenology Flowering May–Jun.
Habitat Sand dunes, ponderosa pine forests, mixed shrublands.
Elevation 1800–2200 m. (5900–7200 ft.)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Penstemon laetus occurs from the Cascade Range of south-central Oregon south through the Sierra Nevada of California to the Western Transverse Ranges of southwestern California. The species comprises three partially sympatric varieties.

The Karuk tribe of northwestern California use an infusion of Penstemon laetus as a psychological aid (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon ammophilus is known from Garfield, Kane, and Washington counties. Populations occur on aeolian sand, derived from the Navajo Sandstone. The fistulose stems of P. ammophilus are unique in Penstemon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx lobes lanceolate, 7–11 × 1–2 mm.
var. leptosepalus
1. Calyx lobes ovate to lanceolate, 4–8.5 × 1–2.8 mm.
→ 2
2. Anthers ovate to elliptic in outline, pollen sacs 1.8–2.3 mm.
var. laetus
2. Anthers sagittate in outline, pollen sacs 2.1–2.8 mm.
var. sagittatus
Source FNA vol. 17, p. 236. FNA vol. 17, p. 161.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. laetus var. laetus, P. laetus var. leptosepalus, P. laetus var. sagittatus
Name authority A. Gray: Proc. Boston Soc. Nat. Hist. 7: 147. (1859) — (as Pentstemon) N. H. Holmgren & L. M. Shultz: Brittonia 34: 381, fig. 1. (1982)
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