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Kral's beardtongue

Utah beardtongue, Utah penstemon

Habit Herbs.
Stems

ascending to erect, 40–70 cm, spreading-puberulent or retrorsely hairy proximally, glandular-pubescent distally, not glaucous.

ascending to erect, 15–60 cm, glaucous.

Leaves

basal and cauline, not leathery, glabrate or puberulent;

basal and proximal cauline 20–70 × (5–)9–20 mm, blade oblanceolate, base tapered, margins entire or ± serrate or crenate, apex obtuse to acute;

cauline 6–9 pairs, sessile, 120–150 × 30–40 mm, blade oblong to lanceolate or ovate-lanceolate, base tapered to clasping, margins serrate to dentate, apex acute to acuminate.

glabrous or obscurely scabrous, especially along margins, glaucous;

basal and proximal cauline 35–80(–100) × 5–20 mm, blade oblanceolate, base tapered, margins entire, apex obtuse to acute;

cauline 2–4 pairs, sessile, 15–75 × 4–26 mm, blade elliptic to lanceolate, rarely ovate, base tapered, margins entire, apex acute.

Thyrses

interrupted, conic, 5–20 cm, axis glandular-pubescent, verticillasters 4–7, cymes 3–7-flowered, 2 per node;

proximal bracts ovate-lanceolate to lanceolate, 16–60(–80) × 3–45 mm, margins entire or serrate;

peduncles and pedicels ascending, glandular-pubescent.

± interrupted, ± secund, 11–25 cm, axis glabrous, verticillasters 5–15, cymes 1–4-flowered;

proximal bracts lanceolate, 6–23 × 1–4 mm;

peduncles and pedicels erect, glabrous.

Flowers

calyx lobes lanceolate, 4–5.5 × 0.8–1.2 mm, glandular-pubescent;

corolla pale lavender to nearly white, with violet nectar guides, tubular-funnelform, 10–16 mm, glandular-pubescent externally, sparsely white-lanate internally abaxially, tube 6–7 mm, throat gradually inflated, 5–7 mm diam., slightly 2-ridged abaxially;

stamens included, pollen sacs opposite, navicular, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 7–8 mm, reaching orifice or slightly exserted, 0.3–0.5 mm diam., tip straight to slightly recurved, distal 4–5 mm moderately to densely pilose, hairs golden, to 1.2 mm;

style 9–12 mm.

calyx lobes ovate, 2.5–4(–5) × 1.5–4(–4.6) mm, margins erose, glabrous;

corolla red to crimson, with red nectar guides, bilaterally symmetric, bilabiate, tubular-salverform, 17–22(–25) mm, glandular-pubescent externally, especially distally, glandular-pubescent internally, tube 6–9 mm, throat slightly inflated, 6–7 mm diam., rounded abaxially;

stamens included, pollen sacs explanate, 0.6–1.1 mm, sutures smooth;

staminode 7–10 mm, flattened distally, 0.4–0.5 mm diam., tip recurved, glabrous, rarely distal 1 mm papillate, papillae golden yellow, to 0.1 mm;

style 10–14 mm.

Capsules

5–6 × 3.5–4 mm, glabrous.

7–10 × 5–7 mm.

Penstemon kralii

Penstemon utahensis

Phenology Flowering May. Flowering Apr–Jun.
Habitat Calcareous soils, juniper-oak-hickory woodlands. Sagebrush shrublands, pinyon-juniper woodlands.
Elevation 200–300 m. (700–1000 ft.) 400–2500 m. (1300–8200 ft.)
Distribution
from FNA
AL; TN
[BONAP county map]
from FNA
AZ; CA; CO; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon kralii is known from the southern Cumberland Plateau. It has been collected in Blount, Jackson, Madison, and Morgan counties, Alabama, and in Franklin County, Tennessee.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon utahensis is widespread in the Colorado River Basin from western Colorado through northwestern Arizona, southeastern California, southern Nevada, and southern Utah.

The Hopi tribe of northeastern Arizona used Penstemon utahensis as a ceremonial and decorative plant (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 210. FNA vol. 17, p. 155.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority D. Estes: J. Bot. Res. Inst. Texas 6: 1, figs. 1, 2. (2012) Eastwood: Zoë 4: 124. (1893)
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