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Steptoe Valley beardtongue, Steptoe Valley penstemon

cut-leaf beardtongue, Richardson's beardtongue, Richardson's penstemon, shrubby penstemon

Habit Subshrubs.
Stems

ascending to erect, 9–30(–35) cm, glabrous.

ascending to erect, 20–95 cm, retrorsely hairy, hairs pointed, not glaucous.

Leaves

basal and cauline, basal not reduced, glabrous;

basal and proximal cauline 15–60(–90) × 8–25(–39) mm, blade oblanceolate to obovate, base tapered, apex rounded to obtuse, sometimes mucronate;

cauline 3–7 pairs, short-petiolate or sessile, (25–)35–80 × 10–20(–28) mm, blade lanceolate to ovate, base tapered to clasping, apex obtuse to acute.

cauline, opposite or subopposite, glabrate or retrorsely hairy, not glaucous;

cauline 5–13 pairs, short-petiolate or sessile, 13–70 × 5–30 mm, blade ovate to lanceolate, base tapered, margins laciniate-dentate to irregularly laciniate-pinnatifid, apex obtuse to acute.

Thyrses

continuous or interrupted, cylindric, (4–)6–20 cm, axis glabrous, verticillasters (3–)5–10(–14), cymes 2–5-flowered;

proximal bracts ovate to lanceolate, (5–)13–56 × (2–)4–24 mm;

peduncles and pedicels glabrous, peduncles to 12 mm.

interrupted, cylindric, 2–25(–48) cm, axis densely glandular-pubescent, verticillasters 3–7(–15), cymes 2–7-flowered, 1 or 2 per node;

proximal bracts lanceolate, 7–43 × 2–16 mm;

peduncles and pedicels spreading, sparsely to densely glandular-pubescent.

Flowers

calyx lobes ovate, 5–8(–12) × 2–3.5 mm, margins entire or erose, broadly scarious, apex acuminate, glabrous;

corolla bluish lavender to lavender or pink, with dark reddish violet or reddish nectar guides, funnelform to ventricose, 15–22 mm, glabrous externally, sparsely white-villous internally abaxially or glabrous, tube 5–7 mm, throat gradually inflated, 5–8 mm diam., rounded abaxially;

stamens included, pollen sacs opposite, (1–)1.2–1.6 mm, sutures papillate;

staminode 8–10 mm, included or reaching orifice, 0.7–1 mm diam., tip strongly recurved, distal 2–5 mm densely villous, hairs yellowish, brownish, or orangish, to 1 mm;

style 10–12 mm.

calyx lobes ovate to lanceolate, 4–7 × 1.3–2.3 mm, glandular-pubescent;

corolla bluish to lavender, or pinkish, with reddish purple nectar guides, ventricose-ampliate, 15–32 mm, glandular-pubescent externally, glabrous internally or sparsely to moderately white-lanate abaxially, tube 7–12 mm, throat gradually inflated, 5–14 mm diam., rounded abaxially;

stamens: longer pair reaching orifice or exserted, filaments glabrous, pollen sacs parallel, 1.3–2 mm, distal 1/2 indehiscent, sides glabrous, sutures denticulate, teeth to 0.2 mm;

staminode 16–23 mm, exserted, 0.3–0.8 mm diam., glabrous or distal 2–8 mm sparsely lanate, hairs yellow, to 2 mm;

style 11–23 mm.

Capsules

8–12 × 4–6 mm.

5–9 × 4–5 mm.

Seeds

2–4 mm.

2n

= 16.

Penstemon immanifestus

Penstemon richardsonii

Phenology Flowering May–Jun.
Habitat Sandy or sandy-loam soils, sagebrush grasslands.
Elevation 1500–2200 m. (4900–7200 ft.)
Distribution
from FNA
NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon immanifestus occurs along the western edge of the Bonneville Basin in eastern Nevada and western Utah, and in the Calcareous Mountains, central Great Basin, and Tonopah regions of eastern and central Nevada. Populations have been documented in Elko, Eureka, Lander, Nye, and White Pine counties, Nevada, and in Juab, Millard, and Tooele counties, Utah.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Penstemon richardsonii consists of three weakly differentiated varieties that are partially sympatric on the eastern slope of the Cascade Range.

The Okanagan-Coville tribe of northeastern Washington and the Paiute tribe of the Great Basin region used Penstemon richardsonii as a drug plant to treat typhoid fever and as a dermatological aid (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas 15–20 mm; styles 11–15 mm; staminodes glabrous or sparsely hairy distally.
var. curtiflorus
1. Corollas 22–32 mm; styles 16–23 mm; staminodes with distal 2–8 mm sparsely lanate, rarely glabrous.
→ 2
2. Leaf blades ovate to lanceolate (in outline), margins laciniate-dentate to irregularly laciniate-pinnatifid.
var. richardsonii
2. Leaf blades ovate, sometimes lanceolate, margins dentate, rarely laciniate-dentate.
var. dentatus
Source FNA vol. 17, p. 120. FNA vol. 17, p. 241.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. richardsonii var. curtiflorus, P. richardsonii var. dentatus, P. richardsonii var. richardsonii
Name authority N. H. Holmgren: Brittonia 30: 334, fig. 1. (1978) Douglas ex Lindley: Bot. Reg. 13: plate 1121. (1828) — (as Pentstemon)
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