low beardtongue, lowly penstemon
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Berry's penstemon, mountain pride, Newberry's beardtongue
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Herbs. |
Subshrubs, cespitose. |
decumbent to erect, (2–)5–30(–65) cm, retrorsely hairy, not glaucous. |
ascending to erect, (5–)10–32 cm, glabrous or retrorsely hairy, not glaucous. |
basal and cauline, not leathery, glabrous or retrorsely hairy; basal and proximal cauline (8–)15–105 × 2–22(–32) mm, blade obovate to ovate or oblanceolate, base tapered, margins entire, sometimes serrulate or serrate, apex obtuse to acute; cauline (1 or)2–4 pairs, sessile or proximals short-petiolate, 10–35(–45) × 1–11 mm, blade obovate to oblanceolate or lanceolate, base tapered to clasping, margins entire, rarely ± serrate, apex obtuse to acute. |
persistent, 4–9 pairs, distals distinctly to slightly smaller than proximals, short-petiolate or sessile, (5–)9–42 × 4–14(–18) mm, blade elliptic to obovate, base tapered, cuneate, or cordate-clasping, margins ± serrulate or serrulate-denticulate to serrate, apex rounded to obtuse or acute, glabrous, glaucous. |
interrupted, cylindric to ± secund, (1–)3–20(–39) cm, axis moderately glandular-pubescent, verticillasters (1–)3–9(–11), cymes (1 or)2–6(–11)-flowered, 2 per node; proximal bracts lanceolate, rarely ovate, 6–34(–48) × 1–11 mm, margins entire; peduncles and pedicels ascending to erect, glandular-pubescent and, usually, retrorsely hairy. |
continuous, ± secund, 2–20 cm, axis glandular-pubescent, verticillasters 4–12, cymes 1-flowered; proximal bracts ovate to lanceolate, 4–11(–20) × 2–8(–15) mm; peduncles and pedicels ascending, glandular-pubescent. |
calyx lobes ovate to lanceolate, 2.8–5(–6) × 1–1.9 mm, margins entire or erose, apex acute, glandular-pubescent; corolla blue to purple or violet, with reddish violet nectar guides, bilabiate, not personate, tubular-funnelform, (7–)8–19 mm, glandular-pubescent externally, sparsely to moderately white- or yellow-lanate internally abaxially, tube 2–5 mm, throat slightly inflated, (3–)4.5–6 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, 0.5–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate; staminode (3–)7–9 mm, included, 0.1–0.3 mm diam., tip slightly recurved, distal 1–4 mm sparsely to densely pilose, hairs yellow or golden yellow, to 0.7 mm (medial hairs sparser and shorter); style 6–12 mm. |
calyx lobes lanceolate, 6–10 × 2–2.5 mm, glandular-pubescent; corolla pink, red, scarlet, lavender, or purple, essentially unlined internally but abaxial ridges sometimes pink to whitish, not personate, funnelform, 22–35 mm, glabrous externally, moderately to densely white-lanate internally abaxially, tube 6–9 mm, throat 5–12 mm diam.; stamens included or longer pair exserted, pollen sacs 0.8–1.7 mm; staminode 8–14 mm, slightly flattened distally, 0.1–0.3 mm diam., tip straight, distal 2–8 mm pubescent to lanate, hairs white or yellowish, to 1 mm; style 20–24 mm. |
4–6 × 2.5–3.5 mm, glabrous. |
(8–)10–13 × (4–)5–8 mm. |
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CA; CO; ID; MT; NV; OR; UT; WA; WY
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CA; NV; OR
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Varieties 3 (3 in the flora). Penstemon humilis is widespread and extremely variable throughout its range in the western United States. Except for two fairly distinct elements in Utah, most variation in the species is contained in the widespread var. humilis. D. D. Keck (1945) and N. H. Holmgren (1984) summarized the ranges and morphologic tendencies of some of the more distinctive phases. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). Penstemon newberryi is known from the Sierra Nevada and Cascade ranges of California, Nevada, and Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Cauline leaf blade margins ± serrate; basal leaves (4–)7–22(–32) mm wide; sw Utah. | var. obtusifolius |
1. Cauline leaf blade margins entire, rarely ± serrate distally; basal leaves 2–20 mm wide; California, nw Colorado, s Idaho, sw Montana, Nevada, e Oregon, Utah, Washington, w Wyoming. | → 2 |
2. Basal leaves puberulent or retrorsely hairy; corollas (7–)12–19 mm; California, nw Colorado, s Idaho, sw Montana, Nevada, e Oregon, Utah, Washington, w Wyoming. | var. humilis |
2. Basal leaves glabrous, sometimes ± puberulent, especially along midveins and margins; corollas 10–13 mm; nc Utah. | var. brevifolius |
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1. Corollas pink to lavender or purple, 27–35 mm; stamens included, pollen sacs 1.2–1.7 mm. | var. berryi |
1. Corollas red, scarlet, or purple, 22–30 mm; stamens: longer pair exserted, pollen sacs 0.8–1.3 mm. | → 2 |
2. Corollas red to scarlet; distal leaves distinctly smaller than proximals. | var. newberryi |
2. Corollas purple; distal leaves slightly smaller than proximals. | var. sonomensis |
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FNA vol. 17, p. 208. |
FNA vol. 17, p. 90. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Dasanthera > sect. Erianthera |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Nuttall ex A. Gray: Proc. Amer. Acad. Arts 6: 69. (1862) — (as Pentstemon) |
A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 6(3): 82, plate 14. (1858) — (as Pentstemon) |
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