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bunchleaf penstemon, foothill beardtongue, foothill penstemon

Habit Subshrubs. Herbs or subshrubs.
Stems

ascending to erect, 30–150 cm, glabrous or retrorsely hairy, hairs pointed, rarely glaucous.

glabrous, puberulent, or retrorsely hairy, rarely glandular-pubescent distally or wholly or retrorsely hairy, glaucous or not.

Leaves

essentially cauline, basal absent or essentially so, opposite, glabrous or retrorsely hairy to puberulent, hairs pointed, rarely glaucous;

cauline 5–11 pairs, petiolate or sessile, 10–100 × 0.5–12 mm, blade oblanceolate to lanceolate or linear, base tapered to clasping, margins entire, apex acute.

basal and cauline, basals sometimes few, or cauline, opposite, sometimes alternate, rarely whorled or subalternate, leathery, rarely not (P. papillatus, P. platyphyllus), glabrous, glabrate, puberulent, retrorsely hairy, canescent, pilose, hirsute, or glandular-pubescent, glaucous or not;

basal and proximal cauline petiolate;

cauline sessile, petiolate, or short-petiolate, blade obovate, ovate, spatulate, oblanceolate, lanceolate, elliptic, oblong, or linear, margins entire or toothed, rarely laciniate-pinnatifid.

Thyrses

interrupted or continuous, cylindric, (6–)10–32 cm, axis glabrous or retrorsely hairy, verticillasters (3–)5–14, cymes 1(–3)-flowered, (1 or)2 per node;

proximal bracts lanceolate to linear, 5–65 × 0.5–5 mm;

peduncles and pedicels ascending to erect, glabrous or retrorsely hairy.

interrupted or continuous, cylindric, sometimes secund, axis glabrous, glandular-pubescent, puberulent, or retrorsely hairy, cymes 1 or 2 per node;

peduncles and pedicels spreading to ascending or erect, rarely appressed.

Flowers

calyx lobes obovate to oblanceolate or ovate to lanceolate, 2.6–6(–7) × 1.3–3.3 mm, glabrous or puberulent;

corolla blue to violet or purple, without nectar guides, ventricose to ventricose-ampliate, 24–38 mm, glabrous externally, glabrous internally, tube 7–10 mm, throat gradually inflated, 8–11 mm diam., 2-ridged abaxially;

stamens included or longer pair reaching orifice or slightly exserted, filaments glabrous, pollen sacs parallel, 2.2–3 mm, distal 1/3–1/2 indehiscent, sides pubescent to lanate, hairs white, to 0.8 mm near filament attachment, sutures denticulate, teeth to 0.2 mm;

staminode 16–19 mm, included, 0.4–0.6 mm diam., glabrous;

style 18–27 mm.

calyx lobes: margins entire or erose, rarely lacerate (P. serrulatus), narrowly scarious or scarious, sometimes herbaceous, rarely broadly scarious (P. serrulatus), glabrous or glandular-pubescent, rarely scabrous, puberulent, pubescent, or ciliolate;

corolla lavender to blue, violet, purple, reddish purple, or pinkish, bilaterally symmetric, ± bilabiate, not personate, funnelform, ventricose, ampliate, or ventricose-ampliate, glabrous or glandular-pubescent externally, glabrous or hairy internally abaxially, throat slightly to gradually inflated, not constricted at orifice, rounded to 2-ridged abaxially;

stamens included or longer pair reaching orifice or exserted, filaments glabrous, rarely hairy distally, pollen sacs parallel, rarely divergent, saccate, dehiscing incompletely, distal 1/5–2/3 indehiscent, connective splitting, sides glabrous or hairy, sutures denticulate, teeth to 0.3 mm;

staminode included to exserted, flattened distally, 0.2–1 mm diam., tip straight, glabrous or distal 5–50(–60)% hairy, hairs to 2.5 mm;

style glabrous.

Capsules

6–9 × 4–5 mm.

glabrous.

Seeds

tan, gray, or brown, angled, rarely patelliform, 0.5–3.4 mm.

2n

= 16, 32.

Penstemon heterophyllus

Penstemon sect. Saccanthera

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
w North America
Discussion

Varieties 3 (3 in the flora).

Penstemon heterophyllus apparently occurs only in California (see discussion under 209c. var. purdyi). The species includes three varieties concentrated in the Coast Ranges from Humboldt County south to San Bernardino and San Diego counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 27 (27 in the flora).

D. D. Keck (1932) included most species of Penstemon with saccate anthers in subg. Saccanthera, sect. Saccanthera. Most members of sect. Saccanthera formed a clade nested in subg. Penstemon in the molecular study by A. D. Wolfe et al. (2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Thyrse axes glabrous; stems glabrous, rarely retrorsely hairy; calyx lobes obovate to oblanceolate, rarely lanceolate.
var. heterophyllus
1. Thyrse axes retrorsely hairy; stems retrorsely hairy; calyx lobes ovate to lanceolate.
→ 2
2. Cauline leaves 15–45 × 0.5–4 mm, axillary fascicles present, sometimes absent.
var. australis
2. Cauline leaves (25–)30–100 × 2–12 mm, axillary fascicles absent, rarely present.
var. purdyi
1. Leaf blade margins serrate, dentate, laciniate-dentate, or laciniate-pinnatifid.
→ 2
2. Stems viscid glandular-pubescent.
P. glandulosus
2. Stems glabrous or retrorsely hairy.
→ 3
3. Calyx lobes glabrous, glabrate, or sparsely puberulent, margins sometimes ciliate.
→ 4
4. Styles 10–16 mm; filaments glabrous.
P. serrulatus
4. Styles 18–25 mm; filaments white-lanate distally, rarely glabrous.
P. venustus
3. Calyx lobes glandular-pubescent.
→ 5
5. Corollas ventricose-ampliate, 15–32 mm; pollen sacs 1.3–2 mm; styles 11–23 mm.
P. richardsonii
5. Corollas funnelform, 13–19 mm; pollen sacs 0.9–1.3 mm; styles 8–12 mm.
→ 6
6. Leaves opposite, sometimes alternate; staminodes: distal 1–4 mm sparsely to moderately lanate, hairs white or yellowish, to 1.5 mm.
P. diphyllus
6. Leaves whorled, sometimes alternate; staminodes: distal 6–7 mm moderately to densely lanate, hairs yellow, to 2.5 mm.
P. triphyllus
1. Leaf blade margins entire or subentire.
→ 7
7. Stems viscid glandular-pubescent.
P. glandulosus
7. Stems glabrous, retrorsely hairy, puberulent, or pilose.
→ 8
8. Staminodes hairy.
→ 9
9. Stems retrorsely hairy, hairs appressed, ± scalelike.
P. tiehmii
9. Stems glabrous, retrorsely hairy, or puberulent, sometimes glabrate or hirsute, hairs pointed.
→ 10
10. Thyrse axes glabrous or retrorsely hairy; corollas glabrous externally or lobe margins sometimes densely white-ciliate.
→ 11
11. Corollas 14–20 mm, glabrous externally; styles 11–14 mm.
P. parvulus
11. Corollas 24–38 mm, glabrous externally except lobe margins densely white-ciliate; styles 18–25 mm.
P. venustus
10. Thyrse axes glandular-pubescent; corollas glandular-pubescent externally.
→ 12
12. Leaves hirsute.
P. scapoides
12. Leaves glabrous, retrorsely hairy, or puberulent, sometimes glabrate.
→ 13
13. Leaves whorled, sometimes alternate, cauline blades linear, sometimes lanceolate.
P. triphyllus
13. Leaves opposite, cauline blades spatulate, oblanceolate, oblong, elliptic, or lanceolate.
→ 14
14. Corollas 15–20(–24) mm; styles 12–15 mm.
P. gracilentus
14. Corollas 24–35 mm; styles 22–30 mm.
→ 15
15. Stems retrorsely hairy or puberulent; pollen sacs: distal 1/3 indehiscent.
P. papillatus
15. Stems glabrous proximally, sparsely glandular-pubescent distally; pollen sacs: distal 1/2–2/3 indehiscent.
P. pudicus
8. Staminodes glabrous.
→ 16
16. Corollas glabrous externally.
→ 17
17. Stems puberulent or retrorsely hairy, at least proximally.
→ 18
18. Styles 11–18 mm; corollas 14–26 mm.
→ 19
19. Pedicels puberulent or retrorsely hairy.
P. cusickii
19. Pedicels glabrous or papillate distally.
P. leonardii
18. Styles 18–27 mm; corollas (20–)22–38 mm.
→ 20
20. Pollen sacs 2.2–3 mm; calyx lobes obovate to oblanceolate or ovate to lanceolate, glabrous or puberulent; staminodes included; California.
P. heterophyllus
20. Pollen sacs 1.5–1.9(–2.1) mm; calyx lobes lanceolate, glabrous; staminodes reaching orifice or exserted; Utah.
P. platyphyllus
17. Stems glabrous.
→ 21
21. Pollen sacs: distal 1/2–2/3 indehiscent, sides glabrous; Nevada, Utah.
→ 22
22. Cauline leaf pairs 2–6; pollen sacs 2.4–2.8 mm; calyx lobes (4–)5–6.3 mm; Nevada.
P. floribundus
22. Cauline leaf pairs 6–14; pollen sacs 1.5–2 mm; calyx lobes 1.8–3.2 mm; Utah.
P. sepalulus
21. Pollen sacs: distal 1/3–1/2 indehiscent, sides hispidulous, pubescent, pilose, or lanate near filament attachments; California, Oregon.
→ 23
23. Corollas 14–20 mm, ventricose; pollen sacs 1.4–1.8 mm; styles 11–14 mm.
P. parvulus
23. Corollas 18–38 mm, ventricose, ventricose-ampliate, or funnelform; pollen sacs 1.8–3.5 mm; styles 16–27 mm.
→ 24
24. Leaves glabrous, glaucous; thyrses secund, axes glabrous; proximal bracts ovate to lanceolate.
P. azureus
24. Leaves glabrous or retrorsely hairy, rarely glaucous; thyrses cylindric, axes glabrous or retrorsely hairy; proximal bracts lanceolate to linear.
P. heterophyllus
16. Corollas glandular-pubescent externally.
→ 25
25. Cauline leaves 0.5–1(–2) mm wide, blades linear; proximal bracts linear.
P. filiformis
25. Cauline leaves (1–)2–24 mm wide, blades ovate to obovate, spatulate, oblanceolate, lanceolate, or linear; proximal bracts ovate to oblanceolate or lanceolate, rarely linear.
→ 26
26. Leaves canescent to pilose.
P. purpusii
26. Leaves glabrous or retrorsely hairy, sometimes glabrate or puberulent.
→ 27
27. Stems retrorsely hairy, hairs appressed, scalelike.
→ 28
28. Cauline leaves 3–7 mm wide, blades oblanceolate to lanceolate, proximals sometimes spatulate; pollen sacs 0.9–1.2 mm.
P. rhizomatosus
28. Cauline leaves 7–24 mm wide, blades ovate to spatulate or lanceolate; pollen sacs 1.1–1.6 mm.
P. tiehmii
27. Stems glabrous, retrorsely hairy, or ± puberulent, hairs pointed.
→ 29
29. Pollen sacs 1–1.5(–1.8) mm, sides glabrous.
→ 30
30. Leaves glabrous, usually glaucous; California.
P. caesius
30. Leaves retrorsely hairy, not glaucous; Nevada, Oregon.
P. kingii
29. Pollen sacs 1.5–3.2 mm, sides hispidulous, pubescent, or pilose to lanate.
→ 31
31. Stems glabrous, sometimes ± puberulent; leaves glabrous, sometimes proximals puberulent, glaucous; pollen sacs 2.4–3.2 mm.
P. neotericus
31. Stems retrorsely hairy; leaves puberulent or retrorsely hairy, sometimes glabrate, not glaucous; pollen sacs 1.5–2.8 mm.
→ 32
32. Corollas 21–35 mm; pollen sacs 1.8–2.8 mm.
P. laetus
32. Corollas 15–25 mm; pollen sacs 1.5–2 mm.
P. roezlii
Source FNA vol. 17, p. 235. FNA vol. 17, p. 229.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera Plantaginaceae > Penstemon > subg. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. heterophyllus var. australis, P. heterophyllus var. heterophyllus, P. heterophyllus var. purdyi
P. azureus, P. caesius, P. cusickii, P. diphyllus, P. filiformis, P. floribundus, P. glandulosus, P. gracilentus, P. heterophyllus, P. kingii, P. laetus, P. leonardii, P. neotericus, P. papillatus, P. parvulus, P. platyphyllus, P. pudicus, P. purpusii, P. rhizomatosus, P. richardsonii, P. roezlii, P. scapoides, P. sepalulus, P. serrulatus, P. tiehmii, P. triphyllus, P. venustus
Synonyms P. subg. saccanthera
Name authority Lindley: Edwards’s Bot. Reg. 22: plate 1899. (1836) — (as Pentstemon heterophyllum) Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 329. (1846) — (as Pentstemon)
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