bunchleaf penstemon, foothill beardtongue, foothill penstemon
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Subshrubs. |
Herbs or subshrubs. |
ascending to erect, 30–150 cm, glabrous or retrorsely hairy, hairs pointed, rarely glaucous. |
glabrous, puberulent, or retrorsely hairy, rarely glandular-pubescent distally or wholly or retrorsely hairy, glaucous or not. |
essentially cauline, basal absent or essentially so, opposite, glabrous or retrorsely hairy to puberulent, hairs pointed, rarely glaucous; cauline 5–11 pairs, petiolate or sessile, 10–100 × 0.5–12 mm, blade oblanceolate to lanceolate or linear, base tapered to clasping, margins entire, apex acute. |
basal and cauline, basals sometimes few, or cauline, opposite, sometimes alternate, rarely whorled or subalternate, leathery, rarely not (P. papillatus, P. platyphyllus), glabrous, glabrate, puberulent, retrorsely hairy, canescent, pilose, hirsute, or glandular-pubescent, glaucous or not; basal and proximal cauline petiolate; cauline sessile, petiolate, or short-petiolate, blade obovate, ovate, spatulate, oblanceolate, lanceolate, elliptic, oblong, or linear, margins entire or toothed, rarely laciniate-pinnatifid. |
interrupted or continuous, cylindric, (6–)10–32 cm, axis glabrous or retrorsely hairy, verticillasters (3–)5–14, cymes 1(–3)-flowered, (1 or)2 per node; proximal bracts lanceolate to linear, 5–65 × 0.5–5 mm; peduncles and pedicels ascending to erect, glabrous or retrorsely hairy. |
interrupted or continuous, cylindric, sometimes secund, axis glabrous, glandular-pubescent, puberulent, or retrorsely hairy, cymes 1 or 2 per node; peduncles and pedicels spreading to ascending or erect, rarely appressed. |
calyx lobes obovate to oblanceolate or ovate to lanceolate, 2.6–6(–7) × 1.3–3.3 mm, glabrous or puberulent; corolla blue to violet or purple, without nectar guides, ventricose to ventricose-ampliate, 24–38 mm, glabrous externally, glabrous internally, tube 7–10 mm, throat gradually inflated, 8–11 mm diam., 2-ridged abaxially; stamens included or longer pair reaching orifice or slightly exserted, filaments glabrous, pollen sacs parallel, 2.2–3 mm, distal 1/3–1/2 indehiscent, sides pubescent to lanate, hairs white, to 0.8 mm near filament attachment, sutures denticulate, teeth to 0.2 mm; staminode 16–19 mm, included, 0.4–0.6 mm diam., glabrous; style 18–27 mm. |
calyx lobes: margins entire or erose, rarely lacerate (P. serrulatus), narrowly scarious or scarious, sometimes herbaceous, rarely broadly scarious (P. serrulatus), glabrous or glandular-pubescent, rarely scabrous, puberulent, pubescent, or ciliolate; corolla lavender to blue, violet, purple, reddish purple, or pinkish, bilaterally symmetric, ± bilabiate, not personate, funnelform, ventricose, ampliate, or ventricose-ampliate, glabrous or glandular-pubescent externally, glabrous or hairy internally abaxially, throat slightly to gradually inflated, not constricted at orifice, rounded to 2-ridged abaxially; stamens included or longer pair reaching orifice or exserted, filaments glabrous, rarely hairy distally, pollen sacs parallel, rarely divergent, saccate, dehiscing incompletely, distal 1/5–2/3 indehiscent, connective splitting, sides glabrous or hairy, sutures denticulate, teeth to 0.3 mm; staminode included to exserted, flattened distally, 0.2–1 mm diam., tip straight, glabrous or distal 5–50(–60)% hairy, hairs to 2.5 mm; style glabrous. |
6–9 × 4–5 mm. |
glabrous. |
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tan, gray, or brown, angled, rarely patelliform, 0.5–3.4 mm. |
= 16, 32. |
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CA
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w North America |
Varieties 3 (3 in the flora). Penstemon heterophyllus apparently occurs only in California (see discussion under 209c. var. purdyi). The species includes three varieties concentrated in the Coast Ranges from Humboldt County south to San Bernardino and San Diego counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 27 (27 in the flora). D. D. Keck (1932) included most species of Penstemon with saccate anthers in subg. Saccanthera, sect. Saccanthera. Most members of sect. Saccanthera formed a clade nested in subg. Penstemon in the molecular study by A. D. Wolfe et al. (2006). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Thyrse axes glabrous; stems glabrous, rarely retrorsely hairy; calyx lobes obovate to oblanceolate, rarely lanceolate. | var. heterophyllus |
1. Thyrse axes retrorsely hairy; stems retrorsely hairy; calyx lobes ovate to lanceolate. | → 2 |
2. Cauline leaves 15–45 × 0.5–4 mm, axillary fascicles present, sometimes absent. | var. australis |
2. Cauline leaves (25–)30–100 × 2–12 mm, axillary fascicles absent, rarely present. | var. purdyi |
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1. Leaf blade margins serrate, dentate, laciniate-dentate, or laciniate-pinnatifid. | → 2 |
2. Stems viscid glandular-pubescent. | P. glandulosus |
2. Stems glabrous or retrorsely hairy. | → 3 |
3. Calyx lobes glabrous, glabrate, or sparsely puberulent, margins sometimes ciliate. | → 4 |
4. Styles 10–16 mm; filaments glabrous. | P. serrulatus |
4. Styles 18–25 mm; filaments white-lanate distally, rarely glabrous. | P. venustus |
3. Calyx lobes glandular-pubescent. | → 5 |
5. Corollas ventricose-ampliate, 15–32 mm; pollen sacs 1.3–2 mm; styles 11–23 mm. | P. richardsonii |
5. Corollas funnelform, 13–19 mm; pollen sacs 0.9–1.3 mm; styles 8–12 mm. | → 6 |
6. Leaves opposite, sometimes alternate; staminodes: distal 1–4 mm sparsely to moderately lanate, hairs white or yellowish, to 1.5 mm. | P. diphyllus |
6. Leaves whorled, sometimes alternate; staminodes: distal 6–7 mm moderately to densely lanate, hairs yellow, to 2.5 mm. | P. triphyllus |
1. Leaf blade margins entire or subentire. | → 7 |
7. Stems viscid glandular-pubescent. | P. glandulosus |
7. Stems glabrous, retrorsely hairy, puberulent, or pilose. | → 8 |
| → 9 |
9. Stems retrorsely hairy, hairs appressed, ± scalelike. | P. tiehmii |
9. Stems glabrous, retrorsely hairy, or puberulent, sometimes glabrate or hirsute, hairs pointed. | → 10 |
10. Thyrse axes glabrous or retrorsely hairy; corollas glabrous externally or lobe margins sometimes densely white-ciliate. | → 11 |
11. Corollas 14–20 mm, glabrous externally; styles 11–14 mm. | P. parvulus |
11. Corollas 24–38 mm, glabrous externally except lobe margins densely white-ciliate; styles 18–25 mm. | P. venustus |
10. Thyrse axes glandular-pubescent; corollas glandular-pubescent externally. | → 12 |
| P. scapoides |
12. Leaves glabrous, retrorsely hairy, or puberulent, sometimes glabrate. | → 13 |
13. Leaves whorled, sometimes alternate, cauline blades linear, sometimes lanceolate. | P. triphyllus |
13. Leaves opposite, cauline blades spatulate, oblanceolate, oblong, elliptic, or lanceolate. | → 14 |
14. Corollas 15–20(–24) mm; styles 12–15 mm. | P. gracilentus |
14. Corollas 24–35 mm; styles 22–30 mm. | → 15 |
15. Stems retrorsely hairy or puberulent; pollen sacs: distal 1/3 indehiscent. | P. papillatus |
15. Stems glabrous proximally, sparsely glandular-pubescent distally; pollen sacs: distal 1/2–2/3 indehiscent. | P. pudicus |
| → 16 |
16. Corollas glabrous externally. | → 17 |
17. Stems puberulent or retrorsely hairy, at least proximally. | → 18 |
18. Styles 11–18 mm; corollas 14–26 mm. | → 19 |
19. Pedicels puberulent or retrorsely hairy. | P. cusickii |
19. Pedicels glabrous or papillate distally. | P. leonardii |
18. Styles 18–27 mm; corollas (20–)22–38 mm. | → 20 |
20. Pollen sacs 2.2–3 mm; calyx lobes obovate to oblanceolate or ovate to lanceolate, glabrous or puberulent; staminodes included; California. | P. heterophyllus |
20. Pollen sacs 1.5–1.9(–2.1) mm; calyx lobes lanceolate, glabrous; staminodes reaching orifice or exserted; Utah. | P. platyphyllus |
| → 21 |
21. Pollen sacs: distal 1/2–2/3 indehiscent, sides glabrous; Nevada, Utah. | → 22 |
22. Cauline leaf pairs 2–6; pollen sacs 2.4–2.8 mm; calyx lobes (4–)5–6.3 mm; Nevada. | P. floribundus |
22. Cauline leaf pairs 6–14; pollen sacs 1.5–2 mm; calyx lobes 1.8–3.2 mm; Utah. | P. sepalulus |
21. Pollen sacs: distal 1/3–1/2 indehiscent, sides hispidulous, pubescent, pilose, or lanate near filament attachments; California, Oregon. | → 23 |
23. Corollas 14–20 mm, ventricose; pollen sacs 1.4–1.8 mm; styles 11–14 mm. | P. parvulus |
23. Corollas 18–38 mm, ventricose, ventricose-ampliate, or funnelform; pollen sacs 1.8–3.5 mm; styles 16–27 mm. | → 24 |
24. Leaves glabrous, glaucous; thyrses secund, axes glabrous; proximal bracts ovate to lanceolate. | P. azureus |
24. Leaves glabrous or retrorsely hairy, rarely glaucous; thyrses cylindric, axes glabrous or retrorsely hairy; proximal bracts lanceolate to linear. | P. heterophyllus |
16. Corollas glandular-pubescent externally. | → 25 |
25. Cauline leaves 0.5–1(–2) mm wide, blades linear; proximal bracts linear. | P. filiformis |
25. Cauline leaves (1–)2–24 mm wide, blades ovate to obovate, spatulate, oblanceolate, lanceolate, or linear; proximal bracts ovate to oblanceolate or lanceolate, rarely linear. | → 26 |
26. Leaves canescent to pilose. | P. purpusii |
26. Leaves glabrous or retrorsely hairy, sometimes glabrate or puberulent. | → 27 |
27. Stems retrorsely hairy, hairs appressed, scalelike. | → 28 |
28. Cauline leaves 3–7 mm wide, blades oblanceolate to lanceolate, proximals sometimes spatulate; pollen sacs 0.9–1.2 mm. | P. rhizomatosus |
28. Cauline leaves 7–24 mm wide, blades ovate to spatulate or lanceolate; pollen sacs 1.1–1.6 mm. | P. tiehmii |
27. Stems glabrous, retrorsely hairy, or ± puberulent, hairs pointed. | → 29 |
29. Pollen sacs 1–1.5(–1.8) mm, sides glabrous. | → 30 |
30. Leaves glabrous, usually glaucous; California. | P. caesius |
30. Leaves retrorsely hairy, not glaucous; Nevada, Oregon. | P. kingii |
29. Pollen sacs 1.5–3.2 mm, sides hispidulous, pubescent, or pilose to lanate. | → 31 |
31. Stems glabrous, sometimes ± puberulent; leaves glabrous, sometimes proximals puberulent, glaucous; pollen sacs 2.4–3.2 mm. | P. neotericus |
31. Stems retrorsely hairy; leaves puberulent or retrorsely hairy, sometimes glabrate, not glaucous; pollen sacs 1.5–2.8 mm. | → 32 |
32. Corollas 21–35 mm; pollen sacs 1.8–2.8 mm. | P. laetus |
32. Corollas 15–25 mm; pollen sacs 1.5–2 mm. | P. roezlii |
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FNA vol. 17, p. 235. |
FNA vol. 17, p. 229. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
Plantaginaceae > Penstemon > subg. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. azureus, P. caesius, P. cusickii, P. diphyllus, P. filiformis, P. floribundus, P. glandulosus, P. gracilentus, P. heterophyllus, P. kingii, P. laetus, P. leonardii, P. neotericus, P. papillatus, P. parvulus, P. platyphyllus, P. pudicus, P. purpusii, P. rhizomatosus, P. richardsonii, P. roezlii, P. scapoides, P. sepalulus, P. serrulatus, P. tiehmii, P. triphyllus, P. venustus |
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P. subg. saccanthera |
Lindley: Edwards’s Bot. Reg. 22: plate 1899. (1836) — (as Pentstemon heterophyllum) |
Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 329. (1846) — (as Pentstemon) |
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