bunchleaf penstemon, foothill beardtongue, foothill penstemon
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stemless beardtongue
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Subshrubs. |
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ascending to erect, 30–150 cm, glabrous or retrorsely hairy, hairs pointed, rarely glaucous. |
prostrate to ascending, to 1 cm (to 15 cm diam.), scabrous or puberulent. |
essentially cauline, basal absent or essentially so, opposite, glabrous or retrorsely hairy to puberulent, hairs pointed, rarely glaucous; cauline 5–11 pairs, petiolate or sessile, 10–100 × 0.5–12 mm, blade oblanceolate to lanceolate or linear, base tapered to clasping, margins entire, apex acute. |
essentially basal, not leathery, scabrous; basal and proximal cauline sessile, 6–15(–22) × 0.6–1.3(–1.5) mm, blade linear, base tapered, margins entire, apex acute. |
interrupted or continuous, cylindric, (6–)10–32 cm, axis glabrous or retrorsely hairy, verticillasters (3–)5–14, cymes 1(–3)-flowered, (1 or)2 per node; proximal bracts lanceolate to linear, 5–65 × 0.5–5 mm; peduncles and pedicels ascending to erect, glabrous or retrorsely hairy. |
essentially absent, verticillasters 1, cymes 1(or 2)-flowered, 1 per node; proximal bracts linear, 5–10 × 0.6–1 mm; peduncles and pedicels glandular-pubescent and scabrous. |
calyx lobes obovate to oblanceolate or ovate to lanceolate, 2.6–6(–7) × 1.3–3.3 mm, glabrous or puberulent; corolla blue to violet or purple, without nectar guides, ventricose to ventricose-ampliate, 24–38 mm, glabrous externally, glabrous internally, tube 7–10 mm, throat gradually inflated, 8–11 mm diam., 2-ridged abaxially; stamens included or longer pair reaching orifice or slightly exserted, filaments glabrous, pollen sacs parallel, 2.2–3 mm, distal 1/3–1/2 indehiscent, sides pubescent to lanate, hairs white, to 0.8 mm near filament attachment, sutures denticulate, teeth to 0.2 mm; staminode 16–19 mm, included, 0.4–0.6 mm diam., glabrous; style 18–27 mm. |
calyx lobes lanceolate, 3.5–5.5 × 0.8–1.5 mm, glandular-pubescent and scabrous; corolla lavender to blue or violet, without nectar guides, funnelform, 11–15 mm, yellowish or white-villous internally abaxially, tube 5–6 mm, throat gradually inflated, not constricted at orifice, 4.5–6 mm diam., slightly 2-ridged abaxially; stamens: longer pair reaching orifice or exserted, pollen sacs widely divergent or opposite, navicular, 0.7–0.9 mm, dehiscing completely, sutures papillate; staminode 7–9 mm, exserted, 0.4–0.5 mm diam., tip straight to slightly recurved, distal 4–7 mm densely pilose, hairs orange, to 0.8 mm; style 7–8 mm. |
6–9 × 4–5 mm. |
2.5–3.5 × 2.5–3 mm. |
= 16, 32. |
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Flowering May–Jul. |
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Semi-barren rock ledges, clayey ridges, gravelly hilltops. |
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1500–2500 m. (4900–8200 ft.) |
CA
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UT; WY |
Varieties 3 (3 in the flora). Penstemon heterophyllus apparently occurs only in California (see discussion under 209c. var. purdyi). The species includes three varieties concentrated in the Coast Ranges from Humboldt County south to San Bernardino and San Diego counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon acaulis is known from the Bridger Basin. Populations are reported from Daggett County, Utah, and Sweetwater County, Wyoming. E. C. Neese (1993) considered populations from Browns Park, Utah, to be transitional between P. acaulis and P. yampaënsis, treating the latter as P. acaulis var. yampaënsis (Penland) Neese. Some specimens from the vicinity of Clay Basin and Red Creek in northwestern Browns Park, Utah, are morphologically intermediate; other specimens from the same area can be assigned to species unambiguously. Given the morphologic distinctness of the majority of specimens, which fall into two basically discrete geographic regions, P. acaulis and P. yampaënsis are here treated as distinct, closely related species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Thyrse axes glabrous; stems glabrous, rarely retrorsely hairy; calyx lobes obovate to oblanceolate, rarely lanceolate. | var. heterophyllus |
1. Thyrse axes retrorsely hairy; stems retrorsely hairy; calyx lobes ovate to lanceolate. | → 2 |
2. Cauline leaves 15–45 × 0.5–4 mm, axillary fascicles present, sometimes absent. | var. australis |
2. Cauline leaves (25–)30–100 × 2–12 mm, axillary fascicles absent, rarely present. | var. purdyi |
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FNA vol. 17, p. 235. |
FNA vol. 17, p. 128. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Lindley: Edwards’s Bot. Reg. 22: plate 1899. (1836) — (as Pentstemon heterophyllum) |
L. O. Williams: Ann. Missouri Bot. Gard. 21: 345. (1934) |
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