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Sierra beardtongue, Sierra penstemon

Mancos penstemon, stiff beardtongue, stiff penstemon

Habit Herbs.
Stems

decumbent, ascending, or erect, 5–65 cm, glabrous or retrorsely hairy, sometimes also glandular-pubescent distally, not glaucous.

ascending to erect, (11–)16–55(–70) cm, glabrous, not glaucous.

Leaves

basal and cauline, not leathery, glabrous;

basal and proximal cauline 5–140 × 3–20 mm, blade spatulate to oblanceolate or elliptic, base tapered, margins entire, apex rounded to obtuse or acute;

cauline 2–5 pairs, short-petiolate or sessile, 10–60(–70) × 1–14 mm, blade oblong to oblanceolate or lanceolate, base tapered to clasping, margins entire, apex obtuse to acute.

basal and cauline, ± leathery or not, glabrous, not glaucous;

basal and proximal cauline 60–90(–150) × 5–12 mm, blade oblanceolate to lanceolate, base tapered, margins entire, apex acute;

cauline 3–5 pairs, sessile, 18–80 × 3–18 mm, blade lanceolate, base tapered to truncate or clasping, apex acute.

Thyrses

interrupted, cylindric, 1–12 cm, axis ± glandular-pubescent, verticillasters 1–6, cymes (1 or)2–6-flowered, 2 per node;

proximal bracts lanceolate, 4–35 × 1–6 mm, margins entire;

peduncles and pedicels erect, ± glandular-pubescent.

continuous or interrupted, ± secund, 5–40 cm, axis glabrous, verticillasters 4–8(–12), cymes 2- or 3(–5)-flowered, 2 per node;

proximal bracts lanceolate, 12–46(–115) × 2–15 mm, apex acuminate;

peduncles and pedicels glabrous or ± glandular, peduncles to 20 mm, pedicels 2–13(–18) mm.

Flowers

calyx lobes oblong to lanceolate, 4–6 × 0.7–1.2 mm, glandular-pubescent;

corolla lavender to blue, violet, or purple, essentially without nectar guides, tubular-funnelform, 10–16 mm, glandular-pubescent externally, moderately yellow-pilose to yellow-lanate internally abaxially, tube 5–7 mm, throat slightly inflated, 3–5 mm diam., prominently 2-ridged abaxially;

stamens: longer pair reaching orifice or slightly exserted, pollen sacs opposite, navicular, 0.6–0.8(–1) mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth;

staminode 6–9 mm, included, 0.3–0.6 mm diam., tip straight to recurved, glabrous or distal 1–3 mm sparsely to moderately pilose, hairs yellow, to 0.5 mm;

style 9–12 mm.

calyx lobes ovate to lanceolate, (4.5–)6–8(–9) × 2–3 mm, apex acuminate to caudate, glabrous or ± glandular proximally;

corolla lavender to blue, without nectar guides, ventricose, (19–)25–32 mm, glabrous externally, glabrous internally, tube 5–7 mm, throat gradually to abruptly inflated, not constricted at orifice, 7–10 mm diam., slightly 2-ridged abaxially;

stamens: longer pair slightly exserted, pollen sacs divergent, navicular-sigmoid, 1.3–2.3 mm, dehiscing incompletely, proximal 1/5–1/4 indehiscent, connective not splitting, sides sparsely to densely lanate, hairs white, to 1 mm, sutures denticulate, teeth to 0.1 mm;

staminode (8–)11–15 mm, included, 0.5–0.1(–1.2) mm diam., tip straight to recurved, distal (2–)4–5 mm sparsely to moderately villous, hairs yellow or white, to 1 mm;

style (11–)13–21 mm.

Capsules

4–6 × 2.5–3.5 mm, glabrous.

7–11 × 4–6 mm.

Penstemon heterodoxus

Penstemon strictiformis

Phenology Flowering May–Jun.
Habitat Juniper and pinyon-juniper woodlands.
Elevation 1700–2400 m. (5600–7900 ft.)
Distribution
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Penstemon heterodoxus comprises three partially sympatric varieties centered in the Sierra Nevada of eastern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants of Penstemon strictiformis from southwestern Colorado (Montezuma County) and northwestern New Mexico (San Juan County) closely match the type, with large corollas, broad calyx lobes, and large pollen sacs. Plants from southeastern Utah have relatively smaller corollas, narrower calyx lobes, and smaller pollen sacs.

Penstemon strictiformis often is confused with P. strictus; many specimens from southwestern Colorado are difficult to place with certainty due to morphologic intermediacy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Thyrse axes usually sparsely glandular-pubescent; n California.
var. shastensis
1. Thyrse axes moderately to densely glandular-pubescent; e California, wc Nevada.
→ 2
2. Stems 5–25 cm; basal and proximal cauline leaves 5–50 × 3–10 mm; verticillasters (1–)3; ec California, wc Nevada.
var. heterodoxus
2. Stems 15–35 cm; basal and proximal cauline leaves (17–)20–75 × 6–18 mm; verticillasters 2–6; se California.
var. cephalophorus
Source FNA vol. 17, p. 207. FNA vol. 17, p. 180.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. heterodoxus var. cephalophorus, P. heterodoxus var. heterodoxus, P. heterodoxus var. shastensis
Synonyms P. strictus subsp. strictiformis
Name authority A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 269. (1878) — (as Pentstemon) Rydberg: Bull. Torrey Bot. Club 31: 642. (1905) — (as Pentstemon)
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