Sierra beardtongue, Sierra penstemon
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low beardtongue, lowly penstemon
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Herbs. |
Herbs. |
decumbent, ascending, or erect, 5–65 cm, glabrous or retrorsely hairy, sometimes also glandular-pubescent distally, not glaucous. |
decumbent to erect, (2–)5–30(–65) cm, retrorsely hairy, not glaucous. |
basal and cauline, not leathery, glabrous; basal and proximal cauline 5–140 × 3–20 mm, blade spatulate to oblanceolate or elliptic, base tapered, margins entire, apex rounded to obtuse or acute; cauline 2–5 pairs, short-petiolate or sessile, 10–60(–70) × 1–14 mm, blade oblong to oblanceolate or lanceolate, base tapered to clasping, margins entire, apex obtuse to acute. |
basal and cauline, not leathery, glabrous or retrorsely hairy; basal and proximal cauline (8–)15–105 × 2–22(–32) mm, blade obovate to ovate or oblanceolate, base tapered, margins entire, sometimes serrulate or serrate, apex obtuse to acute; cauline (1 or)2–4 pairs, sessile or proximals short-petiolate, 10–35(–45) × 1–11 mm, blade obovate to oblanceolate or lanceolate, base tapered to clasping, margins entire, rarely ± serrate, apex obtuse to acute. |
interrupted, cylindric, 1–12 cm, axis ± glandular-pubescent, verticillasters 1–6, cymes (1 or)2–6-flowered, 2 per node; proximal bracts lanceolate, 4–35 × 1–6 mm, margins entire; peduncles and pedicels erect, ± glandular-pubescent. |
interrupted, cylindric to ± secund, (1–)3–20(–39) cm, axis moderately glandular-pubescent, verticillasters (1–)3–9(–11), cymes (1 or)2–6(–11)-flowered, 2 per node; proximal bracts lanceolate, rarely ovate, 6–34(–48) × 1–11 mm, margins entire; peduncles and pedicels ascending to erect, glandular-pubescent and, usually, retrorsely hairy. |
calyx lobes oblong to lanceolate, 4–6 × 0.7–1.2 mm, glandular-pubescent; corolla lavender to blue, violet, or purple, essentially without nectar guides, tubular-funnelform, 10–16 mm, glandular-pubescent externally, moderately yellow-pilose to yellow-lanate internally abaxially, tube 5–7 mm, throat slightly inflated, 3–5 mm diam., prominently 2-ridged abaxially; stamens: longer pair reaching orifice or slightly exserted, pollen sacs opposite, navicular, 0.6–0.8(–1) mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 6–9 mm, included, 0.3–0.6 mm diam., tip straight to recurved, glabrous or distal 1–3 mm sparsely to moderately pilose, hairs yellow, to 0.5 mm; style 9–12 mm. |
calyx lobes ovate to lanceolate, 2.8–5(–6) × 1–1.9 mm, margins entire or erose, apex acute, glandular-pubescent; corolla blue to purple or violet, with reddish violet nectar guides, bilabiate, not personate, tubular-funnelform, (7–)8–19 mm, glandular-pubescent externally, sparsely to moderately white- or yellow-lanate internally abaxially, tube 2–5 mm, throat slightly inflated, (3–)4.5–6 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, 0.5–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate; staminode (3–)7–9 mm, included, 0.1–0.3 mm diam., tip slightly recurved, distal 1–4 mm sparsely to densely pilose, hairs yellow or golden yellow, to 0.7 mm (medial hairs sparser and shorter); style 6–12 mm. |
4–6 × 2.5–3.5 mm, glabrous. |
4–6 × 2.5–3.5 mm, glabrous. |
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CA; NV
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CA; CO; ID; MT; NV; OR; UT; WA; WY
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Varieties 3 (3 in the flora). Penstemon heterodoxus comprises three partially sympatric varieties centered in the Sierra Nevada of eastern California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). Penstemon humilis is widespread and extremely variable throughout its range in the western United States. Except for two fairly distinct elements in Utah, most variation in the species is contained in the widespread var. humilis. D. D. Keck (1945) and N. H. Holmgren (1984) summarized the ranges and morphologic tendencies of some of the more distinctive phases. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Thyrse axes usually sparsely glandular-pubescent; n California. | var. shastensis |
1. Thyrse axes moderately to densely glandular-pubescent; e California, wc Nevada. | → 2 |
2. Stems 5–25 cm; basal and proximal cauline leaves 5–50 × 3–10 mm; verticillasters (1–)3; ec California, wc Nevada. | var. heterodoxus |
2. Stems 15–35 cm; basal and proximal cauline leaves (17–)20–75 × 6–18 mm; verticillasters 2–6; se California. | var. cephalophorus |
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1. Cauline leaf blade margins ± serrate; basal leaves (4–)7–22(–32) mm wide; sw Utah. | var. obtusifolius |
1. Cauline leaf blade margins entire, rarely ± serrate distally; basal leaves 2–20 mm wide; California, nw Colorado, s Idaho, sw Montana, Nevada, e Oregon, Utah, Washington, w Wyoming. | → 2 |
2. Basal leaves puberulent or retrorsely hairy; corollas (7–)12–19 mm; California, nw Colorado, s Idaho, sw Montana, Nevada, e Oregon, Utah, Washington, w Wyoming. | var. humilis |
2. Basal leaves glabrous, sometimes ± puberulent, especially along midveins and margins; corollas 10–13 mm; nc Utah. | var. brevifolius |
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FNA vol. 17, p. 207. |
FNA vol. 17, p. 208. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 269. (1878) — (as Pentstemon) |
Nuttall ex A. Gray: Proc. Amer. Acad. Arts 6: 69. (1862) — (as Pentstemon) |
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