Harbour's beardtongue
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lance-leaf beardtongue
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Herbs. |
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rhizomelike. |
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decumbent, ascending, or weakly erect, 4–18 cm, retrorsely hairy, not glaucous. |
24–75(–98) cm, sparsely to densely retrorsely hairy or puberulent, usually also glandular-pubescent distally. |
cauline, not leathery, glabrate, retrorsely hairy, or puberulent; cauline 2–4 pairs, petiolate or sessile, 7–28 × 3–12 mm, blade spatulate to oblanceolate, base tapered to clasping, margins entire, apex rounded or obtuse to acute. |
retrorsely hairy; basal and proximal cauline 8–80 × 3–12 mm, blade lanceolate or spatulate, base tapered, margins entire, apex rounded to obtuse or acute; cauline 7–15 pairs, 9–85(–110) × 1–11(–22) mm, blade lanceolate to linear, base tapered to truncate, apex acute to acuminate, rarely obtuse. |
interrupted or continuous, ± secund, 1–3 cm, axis ± glandular-pubescent, verticillasters 1 or 2(or 3), cymes 1- or 2-flowered, 1 or 2 per node; proximal bracts lanceolate, 17–23 × 4–10 mm, margins entire; peduncles and pedicels ascending to erect, retrorsely hairy and glandular-pubescent. |
± secund, 5–36 cm, axis glandular-pubescent, verticillasters 3–6, cymes 1 or 2(or 3)-flowered; proximal bracts lanceolate to linear, 5–16 × 1–3 mm; peduncles and pedicels glandular-pubescent, sometimes also retrorsely hairy. |
calyx lobes lanceolate, 6–10 × 1.3–2 mm, glandular-pubescent; corolla lavender or bluish lavender to lavender-purple, without nectar guides, bilabiate, not personate, tubular-funnelform, 15–20 mm, glandular-pubescent externally, densely white-lanate internally abaxially, tube 4–6 mm, throat gradually inflated, 5–6 mm diam., prominently 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate, 0.7–0.9 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 8–9 mm, included, 0.3–0.4 mm diam., tip recurved, distal 4–5 mm moderately to densely lanate, hairs yellow, to 1 mm; style 11–12 mm. |
calyx lobes ovate to lanceolate, 2.9–7 × 1.5–2.5(–3.6) mm, glandular-pubescent; corolla scarlet to red, unlined internally or lined on abaxial surface with faint reddish purple nectar guides, tubular-funnelform, 22–35 mm, glandular-pubescent internally abaxially, tube 5–6 mm, throat gradually inflated, 6–8 mm diam., rounded abaxially; stamens exserted (hidden by galeate adaxial lobes), pollen sacs parallel to divergent, 1.5–2 mm, sutures denticulate, teeth to 0.2 mm; staminode 15–17 mm, included, 0.3–0.4 mm diam.; style 25–30 mm, usually barely exserted from galea. |
6–8 × 4–5 mm, glabrous. |
9–14 × 6–7 mm. |
= 16. |
= 16. |
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Flowering Jun–Aug. |
Flowering May–Sep. |
Alpine talus, gravel slopes, boulder fields. |
Gravelly pinyon-juniper woodlands, pine woodlands, thorn scrub, desert grasslands. |
3200–4200 m. (10500–13800 ft.) |
1200–1800 m. (3900–5900 ft.) |
CO
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AZ; NM; TX; Mexico (Aguascalientes, Chihuahua, Coahuila, Durango, Nayarit, Nuevo León, San Luis Potosí, Tamaulipas, Veracruz, Zacatecas) |
Penstemon harbourii is found on the high peaks of the Central Rocky Mountains. Populations have been documented in at least 17 counties. Habit, pubescence, and root characters of P. harbourii are somewhat anomalous in sect. Penstemon, prompting D. D. Keck (1945) to suggest possible ties with sect. Caespitosi or sect. Cristati. Molecular data appear to provide support for ties with sect. Caespitosi (A. D. Wolfe et al. 2006; C. A. Wessinger et al. 2016). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon lanceolatus is known from southern Arizona (Cochise, Graham, and Greenlee counties), southern New Mexico (Doña Ana, Grant, Hidalgo, Luna, and Sierra counties), and western Texas (Brewster County). Crosswhite published Penstemon ramosus as an avowed substitute for the homonym P. pauciflorus Greene, distinguishing P. ramosus from P. lanceolatus by the former’s branched stems and relatively narrower leaves (1 mm wide versus 4–8 mm wide) with revolute margins. He mapped P. ramosus in southeastern Arizona and southwestern New Mexico and P. lanceolatus in western Texas and northern Mexico. Morphologic differences between these taxa are not consistent enough to warrant recognition of P. ramosus (J. L. Anderson et al. 2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 206. |
FNA vol. 17, p. 109. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Chamaeleon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. ramosus |
A. Gray: Proc. Amer. Acad. Arts 6: 71. (1862) — (as Pentstemon) |
Bentham: Pl. Hartw., 22. (1839) — (as Pentstemon) |
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