Harbour's beardtongue
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Gorman's beardtongue, Gorman's penstemon
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Herbs. |
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rhizomelike. |
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decumbent, ascending, or weakly erect, 4–18 cm, retrorsely hairy, not glaucous. |
ascending to erect, (6–)10–30(–65) cm, glabrous proximally, glandular-pubescent to glandular-villous distally. |
cauline, not leathery, glabrate, retrorsely hairy, or puberulent; cauline 2–4 pairs, petiolate or sessile, 7–28 × 3–12 mm, blade spatulate to oblanceolate, base tapered to clasping, margins entire, apex rounded or obtuse to acute. |
basal and cauline, not leathery, glabrous or distal cauline sometimes glabrous abaxially, glandular-pubescent adaxially; basal and proximal cauline petiolate, (18–)35–100 × (2–)3–10 mm, blade spatulate to oblanceolate, base tapered, margins entire or obscurely serrate, apex rounded to obtuse or acute; cauline 2–4 pairs, sessile, 15–68 × 2–16 mm, blade oblong to lanceolate, base tapered to clasping, margins entire or obscurely serrate, apex obtuse to acute. |
interrupted or continuous, ± secund, 1–3 cm, axis ± glandular-pubescent, verticillasters 1 or 2(or 3), cymes 1- or 2-flowered, 1 or 2 per node; proximal bracts lanceolate, 17–23 × 4–10 mm, margins entire; peduncles and pedicels ascending to erect, retrorsely hairy and glandular-pubescent. |
continuous or ± interrupted, cylindric, (1–)3–15 cm, axis glandular-villous, verticillasters 2–6, cymes (1 or)2–5-flowered, 2 per node; proximal bracts lanceolate to linear, (5–)10–56 × (1–)3–10 mm; peduncles and pedicels glandular-pubescent. |
calyx lobes lanceolate, 6–10 × 1.3–2 mm, glandular-pubescent; corolla lavender or bluish lavender to lavender-purple, without nectar guides, bilabiate, not personate, tubular-funnelform, 15–20 mm, glandular-pubescent externally, densely white-lanate internally abaxially, tube 4–6 mm, throat gradually inflated, 5–6 mm diam., prominently 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate, 0.7–0.9 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 8–9 mm, included, 0.3–0.4 mm diam., tip recurved, distal 4–5 mm moderately to densely lanate, hairs yellow, to 1 mm; style 11–12 mm. |
calyx lobes lanceolate, 7–10.5 × 1.4–2.2 mm, glandular-villous; corolla lavender to purple or bluish violet, with reddish purple nectar guides, ventricose, 19–30 mm, sparsely white- or yellowish villous internally abaxially, tube 5–6 mm, throat abruptly inflated, not constricted at orifice, 8–14 mm diam., rounded abaxially; stamens included, pollen sacs opposite, navicular, 1.3–1.5 mm, dehiscing completely, sutures smooth; staminode 13–17 mm, prominently exserted, 0.4–0.6 mm diam., tip straight to recurved, distal 2–3 mm sparsely lanate, hairs yellow, to 2 mm, and medial 4–8 mm sparsely to moderately pilose, hairs yellow, to 1 mm; style 11–14 mm. |
6–8 × 4–5 mm, glabrous. |
7–10 × 4–5 mm. |
= 16. |
= 16. |
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Flowering Jun–Aug. |
Flowering Jun–Aug. |
Alpine talus, gravel slopes, boulder fields. |
Rocky slopes, sand dunes, gravelly stream terraces, clearings in forests. |
3200–4200 m. (10500–13800 ft.) |
400–1200 m. (1300–3900 ft.) |
CO
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AK; BC; NT; YT |
Penstemon harbourii is found on the high peaks of the Central Rocky Mountains. Populations have been documented in at least 17 counties. Habit, pubescence, and root characters of P. harbourii are somewhat anomalous in sect. Penstemon, prompting D. D. Keck (1945) to suggest possible ties with sect. Caespitosi or sect. Cristati. Molecular data appear to provide support for ties with sect. Caespitosi (A. D. Wolfe et al. 2006; C. A. Wessinger et al. 2016). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon gormanii resembles P. eriantherus vars. argillosus and whitedii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 206. |
FNA vol. 17, p. 136. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
A. Gray: Proc. Amer. Acad. Arts 6: 71. (1862) — (as Pentstemon) |
Greene: Ottawa Naturalist 16: 39. (1902) — (as Pentstemon gormani) |
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