Harbour's beardtongue
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sulphur penstemon, taper-leaf beardtongue, taper-leaf penstemon
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Herbs. |
Herbs. |
rhizomelike. |
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decumbent, ascending, or weakly erect, 4–18 cm, retrorsely hairy, not glaucous. |
ascending to erect, (2–)5–70(–90) cm, glabrous or retrorsely hairy, not glaucous. |
cauline, not leathery, glabrate, retrorsely hairy, or puberulent; cauline 2–4 pairs, petiolate or sessile, 7–28 × 3–12 mm, blade spatulate to oblanceolate, base tapered to clasping, margins entire, apex rounded or obtuse to acute. |
basal and cauline, not leathery, glabrous or puberulent; basal and proximal cauline (30–)55–90(–150) × 5–20(–40) mm, blade spatulate to obovate or elliptic, base tapered, margins entire, rarely denticulate, apex rounded to obtuse or acute; cauline 2–5(–7) pairs, sessile or proximals short- to long-petiolate, 15–90(–130) × 2–25(–38) mm, blade usually lanceolate, proximals sometimes oblanceolate, base tapered to clasping, margins entire, rarely serrate or dentate, apex acute, sometimes obtuse. |
interrupted or continuous, ± secund, 1–3 cm, axis ± glandular-pubescent, verticillasters 1 or 2(or 3), cymes 1- or 2-flowered, 1 or 2 per node; proximal bracts lanceolate, 17–23 × 4–10 mm, margins entire; peduncles and pedicels ascending to erect, retrorsely hairy and glandular-pubescent. |
interrupted, cylindric, 3–27 cm, axis moderately glandular-pubescent, verticillasters 2–7, cymes (1 or)2–8-flowered, 2 per node; proximal bracts lanceolate, 15–55(–80) × 2–15(–28) mm, margins entire or ± serrate to dentate; peduncles and pedicels erect, glandular-pubescent. |
calyx lobes lanceolate, 6–10 × 1.3–2 mm, glandular-pubescent; corolla lavender or bluish lavender to lavender-purple, without nectar guides, bilabiate, not personate, tubular-funnelform, 15–20 mm, glandular-pubescent externally, densely white-lanate internally abaxially, tube 4–6 mm, throat gradually inflated, 5–6 mm diam., prominently 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate, 0.7–0.9 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 8–9 mm, included, 0.3–0.4 mm diam., tip recurved, distal 4–5 mm moderately to densely lanate, hairs yellow, to 1 mm; style 11–12 mm. |
calyx lobes ovate to obovate, ovate-oblong, or lanceolate, 3.2–7.5 × 0.9–2.6 mm, margins erose or lacerate, apex acuminate to short-caudate or long-acuminate, glandular-pubescent; corolla light blue to violet, bluish purple, or pinkish blue, rarely light yellow or whitish, with reddish purple nectar guides, tubular-funnelform, 7–20 mm, glandular-pubescent externally, moderately white-lanate internally abaxially, tube 4–5 mm, throat slightly inflated, 3–5(–6) mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular to slightly saccate, 0.6–1.2 mm, dehiscing completely or incompletely, distal 1/5 or less sometimes indehiscent, connective splitting or not, sides glabrous, sutures smooth or papillate; staminode 7–9 mm, reaching orifice or slightly exserted, 0.3–0.4 mm diam., tip straight to slightly recurved, distal 1–3 mm ± pilose, hairs yellow, to 1.2 mm; style 8–12 mm. |
6–8 × 4–5 mm, glabrous. |
4–8 × 2–5 mm, glabrous. |
= 16. |
= 48. |
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Flowering Jun–Aug. |
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Alpine talus, gravel slopes, boulder fields. |
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3200–4200 m. (10500–13800 ft.) |
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CO
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ID; MT; OR; WA; WY
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Penstemon harbourii is found on the high peaks of the Central Rocky Mountains. Populations have been documented in at least 17 counties. Habit, pubescence, and root characters of P. harbourii are somewhat anomalous in sect. Penstemon, prompting D. D. Keck (1945) to suggest possible ties with sect. Caespitosi or sect. Cristati. Molecular data appear to provide support for ties with sect. Caespitosi (A. D. Wolfe et al. 2006; C. A. Wessinger et al. 2016). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Pollen sacs dehiscing incompletely, distal 1/5 or less usually indehiscent, connectives not splitting, sutures papillate. | var. militaris |
1. Pollen sacs dehiscing completely, connectives splitting, sutures smooth, sometimes ± papillate. | → 2 |
2. Corollas 7–12 mm; calyx lobes lanceolate. | var. palustris |
2. Corollas 12–20 mm; calyx lobes ovate, obovate, ovate-oblong, or lanceolate. | → 3 |
3. Stems (22–)30–70(–90) cm; corollas light blue to violet, rarely light yellow or whitish, 14–20 mm; calyx lobe margins erose. | var. attenuatus |
3. Stems (2–)5–40 cm; corollas bluish purple, 12–16 mm; calyx lobe margins erose or lacerate. | var. pseudoprocerus |
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FNA vol. 17, p. 206. |
FNA vol. 17, p. 193. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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A. Gray: Proc. Amer. Acad. Arts 6: 71. (1862) — (as Pentstemon) |
Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1295. (1830) — (as Pentstemon attenuatum) |
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