Grinnell's beardtongue
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Tushar Range beardtongue
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Herbs. |
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ascending to erect, 10–90 cm, glaucous or not. |
prostrate, spreading, or ascending, 6–12 cm, retrorsely hairy, hairs pointed, sometimes also appressed, white, scalelike. |
basal and proximal cauline 31–90 × 11–20 mm, blade oblanceolate to elliptic, base tapered, margins finely to coarsely dentate, apex rounded to obtuse; cauline 3–6 pairs, short-petiolate or sessile, (15–)30–90 × 4–30 mm, blade lanceolate to ovate-oblong, base tapered, margins coarsely dentate, distals sometimes entire, apex obtuse or acute. |
not leathery, glabrous or sparsely hairy, hairs appressed, white, scalelike abaxially, ± densely hairy, hairs appressed, white, scalelike adaxially; cauline 1–5 pairs, petiolate, 6–16 × 2–4.5 mm, blade obovate to spatulate, base tapered, apex rounded to mucronate or acute. |
interrupted, cylindric, (5–)7–34 cm, axis sparsely to densely glandular-pubescent, verticillasters 4–10, cymes 2–4-flowered; proximal bracts ovate to lanceolate, 5–50 × 2–18 mm; peduncles and pedicels spreading to ascending, glandular-pubescent. |
continuous, secund, 2–12 cm, axis retrorsely hairy, hairs appressed or spreading, white, scalelike, sometimes also glandular-pubescent distally, verticillasters 3–12, cymes 1–3(or 4)-flowered, 1 per node; proximal bracts spatulate to oblanceolate, 8–17 × 1.4–4 mm; peduncles and pedicels spreading to ascending, glandular-pubescent, hairs appressed, white, scalelike. |
calyx lobes ovate, 4.5–8.5 × (2–)2.9–3.2 mm, glandular-pubescent; corolla white to pink or light lavender to light blue or light violet, with reddish purple nectar guides, strongly bilabiate, ventricose-ampliate, 22–35 mm, glandular-pubescent externally, glandular-pubescent internally, also white-villous abaxially, tube 5–8 mm, length 0.9–1.2 times calyx lobes, throat abruptly inflated, constricted at orifice, 10–18 mm diam., rounded abaxially; stamens included, filaments of shorter pair glandular-puberulent proximally, pollen sacs navicular to subexplanate, 1.6–2.2 mm, sutures smooth; staminode 20–22 mm, exserted, 0.4–0.8 mm diam., tip recurved, distal 8–10 mm densely lanate, hairs yellowish or whitish, to 4 mm; proximal 6–8 mm densely glandular-pubescent; style 15–18 mm, glabrous. |
calyx lobes lanceolate to oblong, 3–5 × 0.7–1 mm, herbaceous-margined, glandular-pubescent and retrorsely hairy, hairs appressed, white, scalelike; corolla blue to lavender, or violet, lined internally abaxially with reddish violet nectar guides, ampliate, 12–18 mm, yellow-lanate internally abaxially, tube 5–6 mm, throat gradually inflated, 3.5–5 mm diam., 2-ridged abaxially; stamens reaching orifice or longer pair exserted, filaments glabrous, pollen sacs opposite, navicular to subexplanate, 0.6–0.8 mm, sutures papillate; staminode 7–8 mm, included, flattened distally, 0.2–0.3 mm diam., tips straight to recurved, distal 6–7 mm pubescent, hairs yellow or orange, to 0.7 mm; style 11–14 mm. |
10–14 × 5–8 mm, glabrous. |
3–5 × 3–4 mm. |
= 16. |
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Flowering Apr–Jul. |
Flowering Jun–Aug. |
Chaparral, foothills, pine-juniper woodlands, pine forests. |
Sandy or gravelly slopes, sagebrush shrublands, aspen woodlands, pine woodlands. |
500–2700 m. (1600–8900 ft.) |
2100–3100 m. (6900–10200 ft.) |
CA
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UT |
Two varieties of Penstemon grinnellii have been recognized. Variety grinnellii, with non-glaucous stems 10–60 cm and corollas white to pink and 22–30 mm, has been reported from the Western Transverse and Peninsular ranges, and in the San Bernardino, San Gabriel, and San Jacinto mountains. Variety scrophularioides, with glaucous stems 45–90 cm and corollas light lavender, light blue, or light violet and 26–35 mm, has been reported in the San Francisco Bay Area, southern Sierra Nevada and South Coast and Western Transverse ranges. Characteristics that distinguish the varieties overlap in some populations. Penstemon ×peirsonii Munz & I. M. Johnston, a putative hybrid between P. grinnellii and P. speciosus, and P. ×dubius Davidson, a putative hybrid between P. centranthifolius and P. grinnellii, both have been reported from California (D. D. Keck 1937b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon xylus is known from south-central Utah. Populations are documented in Beaver, Garfield, Iron, Piute, and Sevier counties. The placement of P. xylus remains unclear. D. D. Keck (1937) treated it as P. caespitosus subsp. suffruticosus. N. H. Holmgren (1979b) believed it to be more closely related to P. thompsoniae, which also has vestiture of appressed, white, scalelike hairs. The correct name for Penstemon xylus at the rank of species, which has been a source of confusion, was discussed by C. C. Freeman (2017). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 251. |
FNA vol. 17, p. 107. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. yampaënsis |
P. grinnellii subsp. scrophularioides, P. grinnellii var. scrophularioides, P. scrophularioides |
P. caespitosus var. suffruticosus |
Eastwood: Bull. Torrey Bot. Club 32: 207. (1905) — (as Pentstemon) |
A. Nelson: Bot. Gaz. 34: 31. (1902) — (as Pentstemon) |
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